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Although putative corals of uncertain affinities occur in the early Cambrian, the earliest definite tabulate corals have not been described prior to the Early Ordovician in North America. This paper reports a new finding of a tabulate-like coralomorph forming part of biostratigraphically well-constrained reef mounds in the latest Cambrian–Early Ordovician La Silla Formation in the Argentine Precordillera. The oldest record of the coralomorph genus Amsassia is reported and a new species, A. argentina, is erected. The discovery of this genus in the lowermost Ordovician modifies the previously proposed paleogeographic distribution and patterns of origination and migration routes of this coral-like organism. Amsassia argentina n. sp. constitutes a main framework builder together with a complex microbial consortium. This oldest occurrence of Amsassia as a reef builder represents a new record of a skeletal organism in the gap of metazoan reef constructors after the demise of archaeocyaths in the late early Cambrian.

Trilobites have traditionally been considered fully marine. Through the integration of ichnological, palaeobiological and sedimentological datasets within a sequence-stratigraphic and strati­graphic palaeobiology framework, we challenge this assumption. This analysis is based on the study of incised fluvio-estuarine valley deposits from the Furongian Tilcara Member (TM) and the latest Furongian Pico de Halcón Member (PHM) of the Santa Rosita Formation, the early late Tremadocian Cardonal Formation (CF), and the Dapingian–Darriwilian Alto del Cóndor Formation (ACF), from Cordillera Oriental of northwest Argentina. These valleys were incised into wave-dominated shallow-marine strata and filled with transgressive deposits that accumulated in tide-dominated estuaries. Whereas the TM lacks any body or trace fossil evidence of the presence of trilobites in estuarine settings, the other three units reveal that trilobites were able to inhabit these settings. The PHM and CF are host to trilobite trace fossils in outer estuarine facies, both containing various ichnospecies of Cruziana (e.g., C. omanica and C. semiplicata in the TM) and Rusophycus (e.g., R. latus in both units). In addition, the PHM also contains body fossils of the olenid trilobite Neoparabolina frequens argentina in the same deposits in which the trace fossils are preserved, as well as from middle estuarine facies. The ACF displays trilobite trace fossils of the C. rugosa group in inner, middle, and outer estuarine deposits, illustrating further landward incursions. This unit also contains body fossils of the asaphid trilobite Ogyginus sp. Accordingly, our data indicate two attempts of landward exploration via brackish water: phase 1 in which the outer to middle portion of estuaries were colonized by olenids (Furongian–early late Tremadocian) and phase 2 involving exploration of the inner, middle, and outer estuarine zones by asaphids (Dapingian–Darriwilian). Our study indicates that these trilobites were tolerant to salinity stress and able to make use of the ecological advantages offered by marginal-marine environments migrating up-estuary, following salt wedges either reflecting amphidromy or as euryhaline marine wanderers. It is suggested that tolerance to salinity stress arose independently among different trilobite groups as a result of the broad array of behaviors and adaptations of trilobites during the Ordovician radiation. We speculate that the assumption that all trilobites were stenohaline may have resulted in the misinterpretation of some tide-dominated estuarine deposits as fully marine.

Trilobites, key components of early Palaeozoic communities, are considered to have been invariably fully marine. Through the integration of ichnological, palaeobiological, and sedimentological datasets within a sequence-stratigraphical framework, we challenge this assumption. Here, we report uncontroversial trace and body fossil evidence of their presence in brackish-water settings. Our approach allows tracking of some trilobite groups foraying into tide-dominated estuaries. These trilobites were tolerant to salinity stress and able to make use of the ecological advantages offered by marginal-marine environments migrating up-estuary, following salt wedges either reflecting amphidromy or as euryhaline marine wanderers. Our data indicate two attempts of landward exploration via brackish water: phase 1 in which the outer portion of estuaries were colonized by olenids (Furongian–early late Tremadocian) and phase 2 involving exploration of the inner to middle estuarine zones by asaphids (Dapingian–Darriwilian). This study indicates that tolerance to salinity stress arose independently among different trilobite groups.

Microbialites comprise the mineralized record of early life on Earth and preserve a spectrum of fabrics that reflect complex physical, chemical, and biological interactions. The relatively rarity of microbialites in modern environments, however, challenges our interpretation of ancient structures. Here we report the occurrence of microbial mats, mineral precipitates, and oncoids in the Laguna Negra, a high-altitude hypersaline Andean lake in Catamarca Province, Argentina. Laguna Negra is a Ca-Na-Cl brine where abundant carbonate precipitation takes place. Extreme environmental conditions, including high UV radiation, elevated salinity, and temperature extremes, restrict multicellular life so that mineralization reflects a combination of local hydrologic conditions, lake geochemistry, and microbial activity. The resulting carbonates consist of micritic laminae, botryoidal cement fans, and isopachous cement laminae that are strikingly similar to those observed in Proterozoic stromatolites, providing insight into mechanisms of mineralization. Here, increased saturation with respect to carbonate minerals reflects mixing of spring-fed inlets and lake waters, favoring microbialite formation and preservation. This highlights the importance of hydrological mixing zones in microbialite formation and as taphonomic windows to record microbial activity. Recent discoveries of minerals related to evaporating playa-lake systems on Mars further highlights the potential of Laguna Negra to provide critical insight into biosignature preservation in both terrestrial and extraterrestrial settings.

The Famatina belt, Central Andes, is part of an ancient accretionary margin built along Western Gondwana in the early Palaeozoic. U-Pb ion microprobe analysis of detrital zircons and Sm-Nd whole-rock analysis of two early Palaeozoic low-grade metasedimentary units record the early evolution of this region. Detrital zircons in the Negro Peinado and Achavil formations have ages ranging from Palaeoproterozoic to Cambrian, consistent with derivation from Gondwanan sources. TDM ages suggest that the sedimentary rocks were derived from a composite source area, which separated from the mantle during the Palaeoproterozoic (c. 1.8-1.6 Ga). Constraints from the youngest detrital grains indicate accumulation in a Mid- to Late Cambrian foreland basin adjacent to the inboard Pampean orogenic tract. The dominance of Cambrian ages in the Negro Peinado Formation suggests derivation principally from the eastern Pampean belt whereas the dominance of late Neoproterozoic ages in the Achavil Formation suggests that input from the Pampean belt was overwhelmed by older sources. The paucity of Palaeoproterozoic ages argues against direct input from older areas such as the Rio de la Plata craton. The predominance of Meso- and Neoproterozoic ages over older sources suggests that a Brasiliano-age magmatic arc developed on a Mesoproterozoic basement, probably a southern extension of the Arequipa-Antofalla massif.

The Vinchina Foreland Basin, western Argentina, contains a ∼7 km thick nonmarine stratigraphy, chronologically constrained within the Mio-Pliocene (circa 19-3.4 Ma), and where distribution of Illite/Smectite interstratified phases has shown a progressive smectite-illitization progress (R0 → R1 → R3), is consistent with an incipient burial history. R0 represents randomly mixed-layered illite/smectite normally found at shallow depths, as this ordering is not stable at ∼120C. In the Vinchina Basin, however, the R0 is still persistent at ∼7 km depth, and its appearance even in the deepest levels is consistent with previous interpretations of low burial temperatures based on thermochronologic studies of detrital apatites. The maximum paleotemperature estimation and basin depth imply geothermal gradient as low as ∼15C/km, which allowed an estimate of heat flow values between 33 and 42 mW/m2, that would rise to between ∼40 and 51 mW/m2 when the sedimentation rate (thermal blanketing) is taken into account. These values were only reported for cold basins and represent a paleothermal state of a refrigerated lithosphere. We suggest the central Andes were dominated since the Miocene by heat transfer derived mostly from crustal contributions with a minimum input from the asthenosphere. This refrigerated lithosphere is typical of segments affected by flat subduction. Preliminary thermal models based on previous geodynamic approaches support our conclusions.

The Argentine Precordillera is a continental fragment rifted from the Ouachita embayment of the southern margin of Laurentia (North America) during Cambrian time [about 515 million years ago (Ma)] and accreted to the western margin of Gondwana (South America) during Ordovician time (about 455 Ma). Similarities of Cambrian stratigraphic successions and faunas, Grenville basement rocks, and dimensions link the Argentine Precordillera to the Ouachita embayment. Evidence of rifting during Cambrian time and of a wide ocean basin during Ordovician time indicates that the Precordillera traveled as an independent microcontinent to collide with Gondwana.

The Argentine Precordillera generally is interpreted as a fragment of continental crust and sedimentary cover that was rifted from the Ouachita embayment of southern Laurentia in Cambrian time. Synrift sedimentary successions in the northern part of the Precordillera and in the Birmingham graben of southern Laurentia include thick evaporite deposits. Analyses of strontium isotopes indicate that the evaporites are of late Early Cambrian age, consistent with biostratigraphic correlations. Variations in strontium ratios indicate that these Early Cambrian salinas had a persistent base of marine water and that contributions from meteoric runoff varied through time and with location. The stratigraphic and isotopic data suggest a depositional setting of restricted circulation within a system of extensional fault blocks. The salinas occupied graben blocks that were connected to marine circulation and that intermittently received meteoric runoff carrying fine detritus and radiogenic strontium from horst blocks of continental crust. The evaporites in the Precordillera show less meteoric influence than the evaporites in the Birmingham graben, which was far inboard from the continental margin of southern Laurentia. These relationships are consistent with continental rifting in Early Cambrian time, extension of continental crust, and deposition of synrift sediment in graben blocks that were partially open to marine circulation.

Early Paleozoic paleomagnetic data from NW Argentina and Northern Chile have shown large systematic rotations within two domains: one composed of the Western Puna that yields very large (up to 80°) counter-clockwise rotations, and the other formed by the Famatina Ranges and the Eastern Puna that shows (~40°) clockwise rotations around vertical axes. In several locations, lack of significant rotations in younger rocks constrains this kinematic pattern to have occurred during the Paleozoic. Previous tectonic models have explained these rotations as indicative of rigid-body rotations of large para-autochthonous crustal blocks or terranes. A different but simple tectonic model that accounts for this pattern is presented in which rotations are associated to crustal shortening and tectonic escape due to the collision of the allochthonous terrane of Precordillera in the Late Ordovician. This collision should have generated dextral shear zones in the back arc region of the convergent SW Gondwana margin, where systematic domino-like clockwise rotations of small crustal blocks accommodate crustal shortening. The Western Puna block, bordering the Precordillera terrane to the north, might have rotated counterclockwise as an independent microplate due to tectonic escape processes, in a fashion similar to the present-day relationship between the Anatolia block and the Arabian microplate.

Magnetostratigraphic and paleomagnetic studies on early Andean synorogenic strata (Del Crestón Fm.), in the Famatina Belt (28.7°S, 67.5°W) clarify details of chronology that permit calculation of sedimentation rates within the broken foreland of west Argentina. The Del Crestón Fm represents the first record of broken foreland sedimentation within the southern Central Andean belt and the earliest retroarc volcanic rocks exposed several hundred kilometers from the trench. Twenty-five out of 49 sites collected along the succession presented a primary remanence, as determined through positive fold and reversal tests. Correlation of the local magnetic polarity section with the global polarity time scale indicates that the sedimentation of Del Crestón Fm started at ~16.7 Ma and continued until ~14.5 Ma. The youngest strata are represented by conglomerates bearing abundant Lower Paleozoic granite boulders indicating unroofing of the crystalline basement within the NW Sierras Pampeanas. This result supports the hypothesis of an early broken foreland stage at these latitudes of the Andes, with involvement of the basement in deformation and coeval retroarc volcanism, common attributes of flat-subduction regimes. A mean site paleomagnetic direction of Dec: 6.3°, Inc: -43.6° (α95: 8.0°, N = 24) confirm our earlier intrepretation that the central part of the Famatina Belt within the Sierras Pampeanas did not undergo large vertical axes rotations since the Middle Miocene.