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"Clark, Neil D. L"
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Novel track morphotypes from new tracksites indicate increased Middle Jurassic dinosaur diversity on the Isle of Skye, Scotland
Dinosaur fossils from the Middle Jurassic are rare globally, but the Isle of Skye (Scotland, UK) preserves a varied dinosaur record of abundant trace fossils and rare body fossils from this time. Here we describe two new tracksites from Rubha nam Brathairean (Brothers' Point) near where the first dinosaur footprint in Scotland was found in the 1980s. These sites were formed in subaerially exposed mudstones of the Lealt Shale Formation of the Great Estuarine Group and record a dynamic, subtropical, coastal margin. These tracksites preserve a wide variety of dinosaur track types, including a novel morphotype for Skye: Deltapodus which has a probable stegosaur trackmaker. Additionally, a wide variety of tridactyl tracks shows evidence of multiple theropods of different sizes and possibly hints at the presence of large-bodied ornithopods. Overall, the new tracksites show the dinosaur fauna of Skye is more diverse than previously recognized and give insight into the early evolution of major dinosaur groups whose Middle Jurassic body fossil records are currently sparse.
Journal Article
Small theropod-dominated dinosaur footprint assemblages in the Middle Jurassic Valtos Sandstone and Kilmaluag Formations on the Isle of Skye, Scotland
Middle Jurassic deposits on the Isle of Skye, Scotland, are improving our understanding of the distribution, palaeoenvironmental preferences and behaviour of theropod dinosaurs from a time when the global fossil record is sparse. Here, we describe and classify 185 Bathonian-aged ex situ dinosaur tracks from Skye’s Trotternish Peninsula—many described for the first time and imaged using photogrammetric techniques—into four morphotypes within a new Hebridean series. In the freshwater, closed-lagoonal Kilmaluag Formation at Lùb Score, smaller morphotypes are more abundant than larger equivalents in the freshwater–brackish fluviodeltaic Valtos Sandstone Formation at Valtos. Although assessable outcrops of track-bearing horizons are limited, we infer that the proximity to, or suitability of, specific palaeoenvironments for different-sized trackmakers may influence assemblage composition. Scarce surfaces with multiple tracks indicate potential trackmaker behaviours in respective palaeoenvironments, including foraging at Valtos and post-hatchling care at Lùb Score. The tracks most likely represent traces of a large megalosaurid and multiple smaller-bodied basal coelurosaurian or non-coelurosaurian (e.g. Ceratosauria, Megalosauroidea, Allosauroidea) theropods. The documentation of these trackmakers and their behaviours further enriches our understanding of dinosaur faunas during this poorly known time.
Journal Article
A Comparison of the Dinosaur Communities from the Middle Jurassic of the Cleveland (Yorkshire) and Hebrides (Skye) Basins, Based on Their Ichnites
Despite the Hebrides and Cleveland basins being geographically close, research has not previously been carried out to determine faunal similarities and assess the possibility of links between the dinosaur populations. The palaeogeography of both areas during the Middle Jurassic shows that there were no elevated landmasses being eroded to produce conglomeratic material in the basins at that time. The low-lying landscape and connected shorelines may have provided connectivity between the two dinosaur populations. The dinosaur fauna of the Hebrides and Cleveland basins has been assessed based primarily on the abundant ichnites found in both areas as well as their skeletal remains. In the two basins, the dinosaur faunas are very similar, consisting of non-neosauropod eusauropods, a possible basal titanosauriform, large and small theropods and ornithopods and europodan thyreophorans. The main difference in the faunas is in the sizes. In the Cleveland Basin, the ichnites suggest that there were medium and large theropods alongside small to medium sized ornithopods, whereas, in the Hebrides Basin, the theropods were from small to large and the ornithopods were medium to large. It is suggested that migrations could have taken place between the two areas during the Middle Jurassic. A tentative food chain from the herbivorous dinosaurs to the top predators can be inferred from the footprints.
Journal Article
Caridoid crustaceans from the Ballagan Formation (Tournaisian, Lower Carboniferous) of Willie's Hole, Chirnside, Scottish Borders, UK
The descriptions of two co-occurring cardioid crustaceans from the Ballagan Formation (Tournaisian, Lower Carboniferous) of Chirnside, Scottish Borders, help to resolve the taxonomy of the genus Tealliocaris. Tealliocaris robusta Peach, 1908 is assigned to Schramocaris to form S. robusta (Peach, 1908) comb. nov. on the basis of morphological characters such as the rugosity and position of the branchial carinae as well as the nature of the pleon, and becomes the earliest representative of this genus in Scotland. A new species of Tealliocaris is also recognised from this locality and is described as T. briggsi sp. nov., based on the smooth carapace, lack of pleonic grooves and the number of spines on the scaphocerite and lateral margin of the anterior carapace. The systematic position of the Pendleian specimens identified by Peach (1908) as ‘Tealliocaris robusta var.’ is finally resolved and described as T. weegie sp. nov.
Journal Article
Review of the Dinosaur Remains from the Middle Jurassic of Scotland, UK
Dinosaurs are rare from the Middle Jurassic worldwide. The Isle of Skye, is the only place in Scotland thus far to have produced dinosaur remains. These remains consist mainly of footprints, but also several bones and teeth. These Bajocian and Bathonian remains represent an important collection of a basal eusauropod, early examples of non-neosauropod and possible basal titanosauriform eusauropods, and theropod remains that may belong to an early coelurosaur and a possible megalosaurid, basal tyrannosauroid, or dromaeosaurid. The footprints from here also suggest a rich and diverse dinosaur fauna for which further better diagnosable remains are likely to be found.
Journal Article
Redescription of Americlus rankini (Woodward, 1868) (Pancrustacea: Cyclida: Americlidae) and interpretation of its systematic placement, morphology, and paleoecology
The best-preserved Scottish species of a cyclidan is reinterpreted, based upon recent advances in cyclidan paleobiology. Americlus rankini (Woodward, 1868) is one of the best-preserved members of this crustacean group, and its morphology suggests that it and related forms exhibited a unique and distinct morphology within Multicrustacea. Paleoecological evidence suggests that the animal was most likely free living, possibly as a scavenger, in a marginal marine environment. Americlus lived in Laurentia during the Middle Mississippian (Visean) to Middle Pennsylvanian (Moscovian).
Journal Article
New Information on the Madagascan Middle Jurassic Sauropod Lapparentosaurus madagascariensis
The systematic position of the Middle Jurassic sauropod Lapparentosaurus madagascariensis is not fully understood due to a lack of useful anatomical detail. Despite many new bone fragments from the axial skeleton, post-cranial skeleton, and a hind limb having been previously unearthed, its systematic position has not yet been satisfactorily established. Although this Malagasy taxon is only recognised by two autapomorphies located in the scapula and coracoid, two features of the neural spine, which are reported here, provide additional information on the common autapomorphies shared with the British genus Cetiosaurus. A full description of the femur and neural spine helps to determine some aspects of its relationship to other similar taxa. Remains of Lapparentosaurus madagascariensis have been recovered from mixed facies that may have been deposited in a shallow water lagoon during a transgressive period in the Isallo IIIb subunit in the Majunga Basin.
Journal Article
Phase 1–2 Trial of a SARS-CoV-2 Recombinant Spike Protein Nanoparticle Vaccine
A recombinant SARS-CoV-2 spike protein nanoparticle vaccine delivered in the deltoid muscle on days 0 and 21 was found to be immunogenic at both 5 μg and 25 μg doses. When given with a saponin-based adjuvant, both doses were equally immunogenic, with little or no reactogenicity, and elicited neutralizing antibody titers higher than those in convalescent serum.
Journal Article