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The Indian pangolin (Manis crassicaudata) is under threat due to hunting for local consumption and illegal trafficking of scales and meat. The dearth of scientific studies on the ecology of the M. crassicaudata has impaired accurate assessments of its conservation needs. This study investigated the habitat preference and burrow characteristics of M. crassicaudata in a tropical lowland rainforest in southwest Sri Lanka. A total of 75 burrows (54 feeding burrows and 21 resting burrows) of M. crassicaudata in four different habitat types i.e. secondary forest, Pine-dominated forest, rubber cultivations and tea-dominated home gardens bordering forest were observed using fixed-width transects in order to characterize resting and feeding burrows of this species. The highest density of resting burrows was recorded from the secondary forest (4ha-1), followed by rubber cultivations (2.5ha-1) while no resting burrows were recorded in the Pine-dominated forest and the tea-dominated home gardens bordering forest. Feeding burrows were more abundant in the Pine-dominated forest (5.7ha-1). The burrow depth, burrow opening height, and width were significantly larger in resting burrows compared to feeding burrows. Resting burrows were located at higher elevations (75-100m) with moderately high slopes (450-600), dense canopy cover (>75%) and away from human habitation. Feeding burrows showed a greater variability in terms of associated environmental features. The study further revealed that Indian pangolins exclusively prefer habitats with rocks and boulders under which they dig resting burrows while the location of feeding burrows largely overlaps with the distribution of prey species. The resting burrow design consisted of a bending tunnel that initially slopes downward and then gradually inclines at an angle between 20 and 300, leading to the resting chamber. Our study highlights the importance of conserving fragmented secondary natural forests in changing landscapes of the southwest lowlands of Sri Lanka as these habitats appear to be critical to sustaining populations of M. crassicaudata.

There is a widespread global shift toward renewable energy sources, where the emphasis is on enhancing the utilization of renewable energy due to the rising costs associated with fossil fuels. In this light, biomass pellets made from woody and non-woody biomass and blends have gained increased attention. Extensive research has been conducted globally to enhance the quality of biomass pellets and to explore the potential to combine woody biomass with other non-woody forms of biomass in biomass pellet production. The heterogeneity of the raw materials used and resulting properties of the biomass pellets have led to the establishment of internationally recognized benchmarks such as the International Organization for Standardization (ISO) 17225 standard to regulate pellet quality. In this article, the key mechanical, physical, chemical, and energy properties of pellets made of different non-woody herbaceous biomass are investigated, and the available test values for such properties of the pellets were meta-analyzed. A comparison of the properties of these pellets with the relevant standards was also performed. A meta-analysis of studies on biomass pellet production was conducted via a comprehensive Systematic Literature Review (SLR). The SLR focuses on determining and analyzing the average values for the key physical properties of biomass pellets using woody biomass as a component in concert with other biomass materials. In addition, the optimal range of mixtures of woody and non-woody biomass was reviewed to produce biomass pellets with potential acceptance in the marketplace. The majority of studies included in the SLR concentrate on pellets made from a mixture of biomass materials. The results show that the average values for wood/non-wood mixtures such as pellet diameter, pellet length, moisture content, ash content, fine particle content, gross calorific value, and bulk density were found to adhere to the ISO standards. However, the average mechanical durability fell short of meeting the requirements of the standards. Additional comparisons were nitrogen, sulfur, volatile matter, and fixed carbon content. The findings in this meta-analysis could be useful in directing future research focused on producing high-quality and efficient biomass pellets derived from biomass blends and mixtures.

Promotion of recreational access is an important strategy that helps to impart conservation values to wetlands and aids in fostering sustainable employment for local communities who live in close proximity to wildlife habitats. Unregulated recreational vehicles in coastal wetlands can cause detrimental impacts on coastal avifauna. There is a dearth of information especially in the South Asian context on the impact of recreational disturbance to waterbirds that can inform the management of wetland destinations. This study examined the response of selected water birds to motor vehicles using multispecies experimental disturbance trials conducted at Bundala National Park, Sri Lanka; a Ramsar-listed wetland of global importance. The selected species had significantly different sensitivities to disturbance caused by motor vehicles (log-rank test,χ²= 51.7, P < 0.05). The probability of a bird responding to a moving vehicle parallel to the bird at a distance of 75 m was greater than 0.6 for all selected species (Calidris minuta, Charadrius alexandrinus, Himantopus himantopus, Limosa limosa, Platalea leucorodia and Mycteria leucocephala) except Hydroprogne caspia. Resting birds appeared to be more affected by recreational disturbance when compared to foraging birds. Larger birds (height of adult bird: 80-105 cm) were more sensitive to recreational stimuli, compared to smaller birds (10-20 cm). In general, responses of birds for recreational stimuli, were prominent up to 100 m from the source. Selected waterbirds showed longer response distances in habitats exposed to high vehicle traffic activity suggesting possible habituation of birds to recurring disturbance. Our findings suggest a minimal setback buffer of 100 m needs to be employed to manage recreational disturbance from motor vehicles at Bundala National Park.

The absence of robust species-specific methods to estimate the number of animals in seizures of pangolin scales is a major barrier to effective law enforcement. Therefore, studies focused on developing methods to establish accurate conversion parameters are a priority. This study proposes improved methods to estimate the number of pangolins in the illegal trade to inform law-enforcing authorities. Based on the observations of 25 specimens, Indian pangolins were on average found to possess 511 scales. Three morph-types of scales were identified: broad rhombic (n=411), elongated kite shape (n=69), and folded scales (n=31). The mean dry weight of the three-scale morph-types was 7.5 g, 4.9 g, and 6.2 g. Based on the average frequency and mean dry weight of each scale morph type, the species-specific dry weight of scales for Indian pangolins was 3.6 kg. Accordingly, we propose new and improved methods based on scale morph-type frequencies and species-specific dry weight of scales to estimate the number of Indian pangolins from quantities of scales. Their accuracy was compared with current methods, and the improved methods were found to be more accurate.

One major unanswered question in neuroscience is how the brain transitions between conscious and unconscious states. General anesthetics offer a controllable means to study these transitions. Induction of anesthesia is commonly attributed to drug-induced global modulation of neuronal function, while emergence from anesthesia has been thought to occur passively, paralleling elimination of the anesthetic from its sites in the central nervous system (CNS). If this were true, then CNS anesthetic concentrations on induction and emergence would be indistinguishable. By generating anesthetic dose-response data in both insects and mammals, we demonstrate that the forward and reverse paths through which anesthetic-induced unconsciousness arises and dissipates are not identical. Instead they exhibit hysteresis that is not fully explained by pharmacokinetics as previously thought. Single gene mutations that affect sleep-wake states are shown to collapse or widen anesthetic hysteresis without obvious confounding effects on volatile anesthetic uptake, distribution, or metabolism. We propose a fundamental and biologically conserved concept of neural inertia, a tendency of the CNS to resist behavioral state transitions between conscious and unconscious states. We demonstrate that such a barrier separates wakeful and anesthetized states for multiple anesthetics in both flies and mice, and argue that it contributes to the hysteresis observed when the brain transitions between conscious and unconscious states.

In addition to scoping the impacts of the four most reported sources of recreational disturbance on shorebirds, this study also advances the concept of Tropical Asia (TA) to collectively describe tourist destinations in the ecologically and geopolitically diverse part of the planet that incorporates the tourism megaregion of South and Southeast Asia. At a time of growing global concern about the rapid decline of shorebird populations, many governments in TA are embracing and capitalising on the exponential growth in demand for coastal recreation and tourism across the region. This political response is partly driven by efforts to deliver economic development, aligned to the United Nations Sustainable Development Goals, in order to secure the livelihoods of people living in less developed coastal areas. However, the rapid increase in visitor numbers and the development of infrastructure to support the booming demand for coastal tourism destinations in TA are further exacerbating the pressures on shorebird populations across the region. Despite these growing pressures and the wealth of research reporting on shorebird populations across the Asian flyways, this scoping study identified surprisingly little research that reports on the recreational disturbance (RD) of shorebirds in TA. While undertaken to inform future research, this study also provides a synthesis of management strategies reported in the global literature into a set of management recommendations for coastal destinations in TA.

Camping is a popular activity in the contemporary nature-based tourism domain and rapidly gaining momentum as a key recreational activity in Sri Lanka’s national parks (NPs). Recreational uses such as camping in natural areas can induce significant and often localised resource impacts that can affect soil, vegetation, wildlife and water, with the severity of such impacts varying according to the intensity of use. Hence, monitoring of the biophysical conditions of campsites has become an important component in the reserve management agenda in many places, especially in developed countries. To the best of our knowledge, this is the first study to examine the biophysical impacts associated with the recreation ecology of camping in Sri Lanka. Ten campsites from three dry zone NPs were selected to assess biophysical impacts of camping activities. Field measurements were based on the fixed radial transect method. Gathered data included the total area of the campsite, erosion potential measured as the area of exposed soil (devoid of vegetation or organic litter), number of exposed roots and human damage to trees, number of fireplaces/ fire scars on the ground, visual counts of litter, soil compaction measured by penetrometer, loss of woody debris. This study reports significant levels of environmental degradation related to all the indictors of biophysical impacts at both high and low use campsites. There was no evidence for any difference in the level of environmental degradation associated with high and low use campsites. The loss of natural values associated with campsites negatively impacted visitors’ nature-based experience. These findings highlight the importance of managing biophysical impacts in campsites to provide a high-quality visitor experience, while sustainably managing tourism activities in NPs.

An accurate morphological description and analysis based on reliable data are unavailable for the geographically isolated population of M. crassicaudata in Sri Lanka. This study provides the most updated morphological description of M. crassicaudata with special reference to body measurements directly obtained from 27 specimens collected island-wide. Morphological parameters were recorded under three age classes that were defined based on their body weight (BW) and total body length (TBL); juvenile (BW: <4.3 kg TBL: <56.0 cm), subadult (BW: 4.3–7.3 kg TBL: 56–101 cm), and adult (BW: >7.3 kg TBL: >101 cm) and gender to reveal sexual dimorphism based on morphometric parameters. The TBL of adult males ranged between 137 and 177 cm while body weight ranged between 20.4 and 48.8 kg. The average count of body scales was 511 ± 21. The body scales were found arranged in 13 longitudinal rows with the highest number of scales observed on the vertebral scale row (16 ± 1). Three major scale morphs were identified; broad rhombic scales, elongated kite-shaped scales, and folded shaped scales. Broad rhombic shaped scales was the dominant scale type (80.49%) on the body (405 ± 7). The tail-length to body-length ratio of an Indian pangolin was 0.87. The tail length of an Indian pangolin is a reliable predictor of the TBL and has potential implications in quick field data gathering.

Purpose – This paper demonstrates that the recommendations regarding visitor satisfaction and revisit intention reported in the international literature apply to the management of ecolodges in Sri Lanka. Design/Methodology/Approach – Data from 362 self-report questionnaires completed by visitors between January 2014 and January 2015 were analysed by structural modelling using SPSS and AMOS to confirm the significance that reported direct and indirect relationships of the latent factors ecolodge attributes, tourist motives, visitor satisfaction, and revisit intention have for Sri Lankan ecolodges. Findings – Responses of visitors to Sri Lankan ecolodges were like those of ecolodge visitors in other countries. Ecolodge attributes had a strong direct influence on both international tourist motives to visit Sri Lanka and visitor satisfaction. Further, travel motives and satisfaction have a substantial direct influence on tourist intentions to revisit individual ecolodges and hence Sri Lanka more broadly. Originality of the research – Having confirmed that the factors which influence satisfaction and revisit intention of visitors to Sri Lankan ecolodges are consistent with the research findings from other countries, this is the first study to demonstrate that recommendations from the international ecolodge literature are applicable to and can inform the management and sustainability of ecolodges in Sri Lanka.

 The absence of robust species-specific methods to estimate the number of animals in seizures of pangolin scales is a major barrier to effective law enforcement. Therefore, studies focused on developing methods to establish accurate conversion parameters are a priority. This study proposes improved methods to estimate the number of pangolins in the illegal trade to inform law-enforcing authorities. Based on the observations of 25 specimens, Indian pangolins were on average found to possess 511 scales. Three morph-types of scales were identified: broad rhombic (n=411), elongated kite shape (n=69), and folded scales (n=31). The mean dry weight of the three-scale morph-types was 7.5 g, 4.9 g, and 6.2 g. Based on the average frequency and mean dry weight of each scale morph type, the species-specific dry weight of scales for Indian pangolins was 3.6 kg. Accordingly, we propose new and improved methods based on scale morph-type frequencies and species-specific dry weight of scales to estimate the number of Indian pangolins from quantities of scales. Their accuracy was compared with current methods, and the improved methods were found to be more accurate. Keywords: CITES, conversion parameters, illegal wildlife trade, pangolins, scale seizures, Sri Lanka