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All birds have fundamentally similar patterns of plumage succession. Thus Humphrey and Parkes (1959) proposed a system of nomenclature (the H-P system), based on homologies, that has become standard for molt studies in North America. However, presumably analogous similarities in pattern between first basic and definitive basic plumages have obscured homologies. Many plumages conventionally known as “first basic” are better considered as novel first-cycle plumages that lack homologous counterparts in subsequent cycles. Consequently, current nomenclature does not consistently reflect between-species homologies. Howell and Corben (2000b) proposed that traditional juvenal plumage can be considered an unambiguous starting point for a terminology that better reflects presumed homologies in basic plumages; alternate and other nonbasic plumages may not necessarily be homologous between species. Four underlying strategies of increasing complexity incorporate all known patterns of plumage succession: the Simple Basic Strategy, the Complex Basic Strategy, the Simple Alternate Strategy, and the Complex Alternate Strategy. We review inconsistency in the H-P system; explain the four underlying strategies; and discuss how one can identify homologies (if any) between plumages in first and subsequent cycles and among taxa. Many species have novel plumages added into their first plumage cycle; we argue that existing terminology for these plumages is unsuitable and we term them formative plumages attained by preformative molts. Finally, we provide examples of how this modified H-P system can be applied to diverse taxa of birds while reflecting the homology underlying all basic plumage cycles. Our revision validates the flexibility and utility of the H-P system. El Problema del Primer Plumaje Básico: Una Revisión de las Homologías de la Muda y del Plumaje Resumen. Todas las aves tienen patrones de sucesión del plumaje fundamentalmente similares. De este modo, Humphrey y Parkes (1959) propusieron un sistema de nomenclatura (el sistema H-P), basado en homologías, el cual ha sido de uso común en estudios de muda de plumaje en Norte América. Sin embargo, supuestas similitudes análogas entre el primer plumaje básico y el plumaje definitivo básico han confundido las homologías. Muchos plumajes convencionalmente conocidos como “primer básico” son considerados mejor como plumajes originales del primer ciclo que carecen de contrapartes homólogas en los ciclos siguientes. Consecuentemente, la nomenclatura actual no refleja las homologías entre especies. Howell y Corben (2000b) propusieron que el tradicional plumaje juvenil puede ser considerado como un punto de partida inequívoco para una terminología que refleje mejor las homologías presuntas en los plumajes básicos; los plumajes alternos y otros plumajes no básicos pudieran no ser homólogos entre especies. Cuatro estrategias de creciente complejidad incorporan todos los patrones conocidos de sucesión de plumajes: La Estrategia Básica Simple, La Estrategia Básica Compleja, La Estrategia Alterna Simple, y La Estrategia Alterna Compleja. Examinamos ciertas inconsistencias en el sistema H-P; explicamos las cuatro estrategias subyacentes, y discutimos cómo se pueden identificar homologías (cuando existen) entre los plumajes del primer ciclo y de los ciclos siguientes, y entre taxa diferentes. Muchas especies tienen plumajes originales adicionales en su primer ciclo de plumaje; sostenemos que la terminología actual para estos plumajes es inadecuada y los denominamos como plumajes formativos, logrados por mudas preformativas. Finalmente, damos ejemplos de como este sistema H-P modificado puede ser aplicado a diversos tipos de aves y al mismo tiempo reflejar la homología subyacente a todos los ciclos de plumajes básicos. Nuestra revisión valida la flexibilidad y utilidad del sistema H-P.

Rare Birds of North Americais the first comprehensive illustrated guide to the vagrant birds that occur throughout the United States and Canada. Featuring 275 stunning color plates, this book covers 262 species originating from three very different regions--the Old World, the New World tropics, and the world's oceans. It explains the causes of avian vagrancy and breaks down patterns of occurrence by region and season, enabling readers to see where, when, and why each species occurs in North America. Detailed species accounts describe key identification features, taxonomy, age, sex, distribution, and status. Rare Birds of North Americaprovides unparalleled insights into vagrancy and avian migration, and will enrich the birding experience of anyone interested in finding and observing rare birds. Covers 262 species of vagrant birds found in the United States and CanadaFeatures 275 stunning color plates that depict every speciesExplains patterns of occurrence by region and seasonProvides an invaluable overview of vagrancy patterns and migrationIncludes detailed species accounts and cutting-edge identification tips

This guide, designed for quick use on day trips off the East Coast, helps you put a name to what you find, from whales and dolphins to shearwaters, turtles, and even flying fish.

Petrels, albatrosses, and storm-petrels are among the most beautiful yet least known of all the world's birds, living their lives at sea far from the sight of most people. Largely colored in shades of gray, black, and white, these enigmatic and fast-flying seabirds can be hard to differentiate, particularly from a moving boat. Useful worldwide, not just in North America, this photographic guide is based on unrivaled field experience and combines insightful text and hundreds of full-color images to help you identify these remarkable birds. The first book of its kind, this guide features an introduction that explains ocean habitats and the latest developments in taxonomy. Detailed species accounts describe key identification features such as flight manner, plumage variation related to age and molt, seasonal occurrence patterns, and migration routes. Species accounts are arranged into groups helpful for field identification, and an overview of unique identification challenges is provided for each group. The guide also includes distribution maps for regularly occurring species as well as a bibliography, glossary, and appendixes. The first state-of-the-art photographic guide to these enigmatic seabirdsIncludes hundreds of full-color photos throughoutFeatures detailed species accounts that describe flight, plumage, distribution, and moreProvides overviews of ocean habitats, taxonomy, and conservationOffers tips on how to observe and identify birds at sea

\"Two-thirds of our planet lies out of sight of land, just offshore beyond the horizon. What wildlife might you find out there? And how might you identify what you see? This Offshore Sea Life ID Guide, designed for quick use on day trips off the West Coast, helps you put a name to what you see, from whales and dolphins to albatrosses, turtles, and even flyingfish. Carefully crafted color plates show species as they typically appear at sea, and expert text highlights identification features. This user-friendly field guide is essential for anyone going out on a whale-watching or birding trip, and provides a handy gateway to the wonders of the ocean.\"--Publisher description.

Ornithologists have largely embraced the molt terminology of Humphrey and Parkes (1959) as modified by Howell et al. (2003; the H-P-H system). In a recent commentary, Wolfe et al. (2014) summarized the derivation and benefits of H-P-H terminology, suggested slight modifications, and promoted analyses on the evolution of molts using H-P-H nomenclature. We appreciate the timeliness of Wolfe et al.'s review and agree with most of their conclusions and modifications. We disagree, however, with Wolfe et al.'s proposal for introducing a new and restricted use of the term “definitive” in H-P-H nomenclature. To avoid confusion, we recommend that definitive plumage and definitive molt cycle continue to be used as defined by Humphrey and Parkes (1959) and Howell et al. (2003), respectively, as terms indicating that plumage appearance and molt cycle have achieved stasis. We also recommend that the term “plumage” can be used more widely than the definition proposed by Humphrey and Parkes (1959), and that the term “juvenal” can henceforth be replaced by “juvenile” in molt and plumage literature.

We document the molt sequence of flight feathers in Turkey Vultures (Cathartes aura) based on studies of captive and wild birds, and examination of museum specimens. We found an unusual pattern of primary replacement, which appears to be a modified form of Staffelmauser, or stepwise wing molt. A Staffelmauser-like strategy for replacement of the secondaries is also described. These patterns of feather replacement appear to be adaptations to maintain flying performance while replacing all primaries and most secondaries during each molt. To what extent molt patterns in Turkey Vultures reflect convergent adaptation for flight, rather than ancestral characters useful for phylogenetic studies, remains unknown.