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The apple ( [Suckow] Borkh.) is one of the most economically and culturally significant fruits in the world today, and it is grown in all temperate zones. With over a thousand landraces recognized, the modern apple provides a unique case study for understanding plant evolution under human cultivation. Recent genomic and archaeobotanical studies have illuminated parts of the process of domestication in the Rosaceae family. Interestingly, these data seem to suggest that rosaceous arboreal crops did not follow the same pathway toward domestication as other domesticated, especially annual, plants. Unlike in cereal crops, tree domestication appears to have been rapid and driven by hybridization. Apple domestication also calls into question the concept of centers of domestication and human intentionality. Studies of arboreal domestication also illustrate the importance of fully understanding the seed dispersal processes in the wild progenitors when studying crop origins. Large fruits in Rosaceae evolved as a seed-dispersal adaptation recruiting megafaunal mammals of the late Miocene. Genetic studies illustrate that the increase in fruit size and changes in morphology during evolution in the wild resulted from hybridization events and were selected for by large seed dispersers. Humans over the past three millennia have fixed larger-fruiting hybrids through grafting and cloning. Ultimately, the process of evolution under human cultivation parallels the natural evolution of larger fruits in the clade as an adaptive strategy, which resulted in mutualism with large mammalian seed dispersers (disperser recruitment).

Wheat and barley evolved from large-seeded annual grasses in the arid, low latitudes of Asia; their spread into higher elevations and northern latitudes involved corresponding evolutionary adaptations in these plants, including traits for frost tolerance and shifts in photoperiod sensitivity. The adaptation of farming populations to these northern latitudes was also a complex and poorly understood process that included changes in cultivation practices and the varieties of crops grown. In this article, we push back the earliest dates for the spread of wheat and barley into northern regions of Asia as well as providing earlier cultural links between East and West Asia. The archaeobotanical, palynological and anthracological data we present come from the Tongtian Cave site in the Altai Mountains, with a punctuated occupation dating between 5,200 and 3,200 calibrated years  bp , coinciding with global cooling of the middle–late Holocene transition. These early low-investment agropastoral populations in the north steppe area played a major role in the prehistoric trans-Eurasian exchange. The spread of wheat and barley involves evolutionary and human adaptation but has been difficult to date properly. Findings from a cave site suggest a much earlier transfer of cereal grains across Eurasia.

The oasis villages of the Tarim Basin served as hubs along the ancient Silk Road, and they played an important role in facilitating communication between the imperial centers of Asia. These villages were supported by an irrigated form of cereal farming that was specifically adapted to these early oasis settlements. In this manuscript, we present the results from new archaeobotanical analyses, radiocarbon dating, and organic carbon isotopic studies directly from carbonized seeds at the Wupaer site (1500–400 BC) in the Kashgar Oasis of the western Tarim Basin. Our results showed that early farming in the oasis relied on a mixed wheat and barley system, but after 1200 BC was intensified through more elaborate irrigation, the introduction of more water-demanding legumes, and possibly a greater reliance on free-threshing wheat. These crops and the knowledge of irrigated farming likely dispersed into the Tarim Basin through the mountains from southern Central Asia. Improved agricultural productivity in the Tarim Basin may also have led to demographic and socio-political shifts and fed into the increased exchange that is colloquially referred to as the Silk Road.

While classic models for the emergence of pastoral groups in Inner Asia describe mounted, horse-borne herders sweeping across the Eurasian Steppes during the Early or Middle Bronze Age (ca. 3000–1500 BCE), the actual economic basis of many early pastoral societies in the region is poorly characterized. In this paper, we use collagen mass fingerprinting and ancient DNA analysis of some of the first stratified and directly dated archaeofaunal assemblages from Mongolia’s early pastoral cultures to undertake species identifications of this rare and highly fragmented material. Our results provide evidence for livestock-based, herding subsistence in Mongolia during the late 3rd and early 2nd millennia BCE. We observe no evidence for dietary exploitation of horses prior to the late Bronze Age, ca. 1200 BCE – at which point horses come to dominate ritual assemblages, play a key role in pastoral diets, and greatly influence pastoral mobility. In combination with the broader archaeofaunal record of Inner Asia, our analysis supports models for widespread changes in herding ecology linked to the innovation of horseback riding in Central Asia in the final 2nd millennium BCE. Such a framework can explain key broad-scale patterns in the movement of people, ideas, and material culture in Eurasian prehistory.

An amendment to this paper has been published and can be accessed via a link at the top of the paper.An amendment to this paper has been published and can be accessed via a link at the top of the paper.