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In the southern winter polar stratosphere, the distribution of gravity wave momentum flux in many state-of-the-art climate simulations is inconsistent with long-time satellite and superpressure balloon observations around 60∘ S. Recent studies hint that a lateral shift between prominent gravity wave sources in the tropospheric mid-latitudes and the location where gravity wave activity is present in the stratosphere causes at least part of the discrepancy. This lateral shift cannot be represented by the column-based gravity wave drag parameterisations used in most general circulation models. However, recent high-resolution analysis and re-analysis products of the European Centre for Medium-Range Weather Forecasts Integrated Forecast System (ECMWF-IFS) show good agreement with the observations and allow for a detailed investigation of resolved gravity waves, their sources, and propagation paths. In this paper, we identify resolved gravity waves in the ECMWF-IFS analyses for a case of high gravity wave activity in the lower stratosphere using small-volume sinusoidal fits to characterise these gravity waves. The 3D wave vector together with perturbation amplitudes, wave frequency, and a fully described background atmosphere are then used to initialise the Gravity Wave Regional or Global Ray Tracer (GROGRAT) gravity wave ray tracer and follow the gravity waves backwards from the stratosphere. Finally, we check for the indication of source processes on the path of each ray and, thus, quantitatively attribute gravity waves to sources that are represented within the model. We find that stratospheric gravity waves are indeed subject to far (>1000 km) lateral displacement from their sources, which already take place at low altitudes (<20 km). Various source processes can be linked to waves within stratospheric gravity wave (GW) patterns, such as the orography equatorward of 50∘ S and non-orographic sources above the Southern Ocean. These findings may explain why superpressure balloons observe enhanced gravity wave momentum fluxes in the lower stratosphere over the Southern Ocean despite an apparent lack of sources at this latitude. Our results also support the need to improve gravity wave parameterisations to account for meridional propagation.

Large uncertainties remain with respect to the representation of atmospheric gravity waves (GWs) in general circulation models (GCMs) with coarse grids. Insufficient parameterizations result from a lack of observational constraints on the parameters used in GW parameterizations as well as from physical inconsistencies between parameterizations and reality. For instance, parameterizations make oversimplifying assumptions about the generation and propagation of GWs. Increasing computational capabilities now allow GCMs to run at grid spacings that are sufficiently fine to resolve a major fraction of the GW spectrum. This study presents the first intercomparison of resolved GW pseudomomentum fluxes (GWMFs) in global convection-permitting simulations and those derived from satellite observations. Six simulations of three different GCMs are analyzed over the period of one month of August to assess the sensitivity of GWMF to model formulation and horizontal grid spacing. The simulations reproduce detailed observed features of the global GWMF distribution, which can be attributed to realistic GWs from convection, orography, and storm tracks. Yet the GWMF magnitudes differ substantially between simulations. Differences in the strength of convection may help explain differences in the GWMF between simulations of the same model in the summer low latitudes where convection is the primary source. Across models, there is no evidence for a systematic change with resolution. Instead, GWMF is strongly affected by model formulation. The results imply that validating the realism of simulated GWs across the entire resolved spectrum will remain a difficult challenge not least because of a lack of appropriate observational data.

Gravity waves are important drivers of dynamic processes in particular in the middle atmosphere. To analyse atmospheric data for gravity wave signals, it is essential to separate gravity wave perturbations from atmospheric variability due to other dynamic processes. Common methods to separate small-scale gravity wave signals from a large-scale background are separation methods depending on filters in either the horizontal or vertical wavelength domain. However, gravity waves are not the only process that could lead to small-scale perturbations in the atmosphere. Recently, concerns have been raised that vertical wavelength filtering can lead to misinterpretation of other wave-like perturbations, such as inertial instability effects, as gravity wave perturbations. In this paper we assess the ability of different spectral background removal approaches to separate gravity waves and inertial instabilities using artificial inertial instability perturbations, global model data and satellite observations. We investigate a horizontal background removal (which applies a zonal wavenumber filter with additional smoothing of the spectral components in meridional and vertical direction), a sophisticated filter based on 2D time–longitude spectral analysis (see Ern et al., 2011) and a vertical wavelength Butterworth filter. Critical thresholds for the vertical wavelength and zonal wavenumber are analysed. Vertical filtering has to cut deep into the gravity wave spectrum in order to remove inertial instability remnants from the perturbations (down to 6 km cutoff wavelength). Horizontal filtering, however, removes inertial instability remnants in global model data at wavenumbers far lower than the typical gravity wave scales for the case we investigated. Specifically, a cutoff zonal wavenumber of 6 in the stratosphere is sufficient to eliminate inertial instability structures. Furthermore, we show that for infrared limb-sounding satellite profiles it is possible as well to effectively separate perturbations of inertial instabilities from those of gravity waves using a cutoff zonal wavenumber of 6. We generalize the findings of our case study by examining a 1-year time series of SABER (Sounding of the Atmosphere using Broadband Emission Radiometry) data.

Atmospheric gravity waves (GWs) are an important coupling mechanism in the middle atmosphere. For instance, they provide a large part of the driving of long-period atmospheric oscillations such as the Quasi-Biennial Oscillation (QBO) and the semiannual oscillation (SAO) and are in turn modulated. They also induce the wind reversal in the mesosphere–lower thermosphere region (MLT) and the residual mean circulation at these altitudes. In this study, the variations in monthly zonal mean gravity wave square temperature amplitudes (GWSTAs) and, for the first time, absolute gravity wave momentum flux (GWMF) on different timescales such as the annual, semiannual, terannual and quasi-biennial variations are investigated by spectrally analyzing SABER observations from 2002 to 2015. Latitude–altitude cross sections of spectral amplitudes and phases of GWSTA and absolute GWMF in the stratosphere and mesosphere are presented and physically interpreted. It is shown that the time series of GWSTA and GWMF at a certain altitude and latitude results from the complex interplay of GW sources, propagation through and filtering in lower altitudes, oblique propagation superposing GWs from different source locations, and, finally, the modulation of the GW spectrum by the winds at a considered altitude and latitude. The strongest component is the annual variation, dominated in the summer hemisphere by subtropical convective sources and in the winter hemisphere by polar vortex dynamics. At heights of the wind reversal, a 180∘ phase shift also occurs, which is at different altitudes for GWSTA and GWMF. In the intermediate latitudes a semiannual variation (SAV) is found. Dedicated GW modeling is used to investigate the nature of this SAV, which is a different phenomenon from the tropical SAO also seen in the data. In the tropics a stratospheric and a mesospheric QBO are found, which are, as expected, in antiphase. Indication for a QBO influence is also found at higher latitudes. In previous studies a terannual variation (TAV) was identified. In the current study we explain its origin. In particular the observed patterns for the shorter periods, SAV and TAV, can only be explained by poleward propagation of GWs from the lower-stratosphere subtropics into the midlatitude and high-latitude mesosphere. In this way, critical wind filtering in the lowermost stratosphere is avoided and this oblique propagation is hence likely an important factor for MLT dynamics.

Many gravity wave analyses, based on either observations or model simulations, assume the presence of only a single dominant wave. This paper shows that there are much more complex cases with gravity waves from multiple sources crossing each others' paths. A complex gravity wave structure consisting of a superposition of multiple wave packets was observed above southern Scandinavia on 28 January 2016 with the Gimballed Limb Observer for Radiance Imaging of the Atmosphere (GLORIA). The tomographic measurement capability of GLORIA enabled a detailed 3-D reconstruction of the gravity wave field and the identification of multiple wave packets with different horizontal and vertical scales. The larger-scale gravity waves with horizontal wavelengths of around 400 km could be characterised using a 3-D wave-decomposition method. The smaller-scale wave components with horizontal wavelengths below 200 km were discussed by visual inspection. For the larger-scale gravity wave components, a combination of gravity-wave ray-tracing calculations and ERA5 reanalysis fields identified orography as well as a jet-exit region and a low-pressure system as possible sources. All gravity waves are found to propagate upward into the middle stratosphere, but only the orographic waves stay directly above their source. The comparison with ERA5 also shows that ray tracing provides reasonable results even for such complex cases with multiple overlapping wave packets. Despite their coarser vertical resolution compared to GLORIA measurements, co-located AIRS measurements in the middle stratosphere are in good agreement with the ray tracing and ERA5 results, proving once more the validity of simple ray-tracing models. Thus, this paper demonstrates that the high-resolution GLORIA observations in combination with simple ray-tracing calculations can provide an important source of information for enhancing our understanding of gravity wave propagation.

Atmospheric gravity waves are a major cause of uncertainty in atmosphere general circulation models. This uncertainty affects regional climate projections and seasonal weather predictions. Improving the representation of gravity waves in general circulation models is therefore of primary interest. In this regard, measurements providing an accurate 3-D characterization of gravity waves are needed. Using the Gimballed Limb Observer for Radiance Imaging of the Atmosphere (GLORIA), the first airborne implementation of a novel infrared limb imaging technique, a gravity wave event over Iceland was observed. An air volume disturbed by this gravity wave was investigated from different angles by encircling the volume with a closed flight pattern. Using a tomographic retrieval approach, the measurements of this air mass at different angles allowed for a 3-D reconstruction of the temperature and trace gas structure. The temperature measurements were used to derive gravity wave amplitudes, 3-D wave vectors, and direction-resolved momentum fluxes. These parameters facilitated the backtracing of the waves to their sources on the southern coast of Iceland. Two wave packets are distinguished, one stemming from the main mountain ridge in the south of Iceland and the other from the smaller mountains in the north. The total area-integrated fluxes of these two wave packets are determined. Forward ray tracing reveals that the waves propagate laterally more than 2000 km away from their source region. A comparison of a 3-D ray-tracing version to solely column-based propagation showed that lateral propagation can help the waves to avoid critical layers and propagate to higher altitudes. Thus, the implementation of oblique gravity wave propagation into general circulation models may improve their predictive skills.

Background Parafilaria bovicola (Nematoda: Filariidae) causes cutaneous bleedings in bovine species. Flies serve as intermediate hosts. In recent years, reports on bovine parafilariosis have become more frequent, corroborating the necessity of reliable diagnostic interventions especially since no molecular or serological test has been available. We aimed to establish a polymerase chain reaction assay to detect DNA of P. bovicola in flies, skin biopsies and serohemorraghic exudates of bleeding spots. Methods PCRs targeting the cytochrome c oxidase subunit 1 ( cox 1) gene and the internal transcribed spacer region (ITS) of the ribosomal RNA gene cluster were evaluated for their diagnostic sensitivity as well as performance and specificity on biopsy and serohemorrhagic exudate samples from P. bovicola- infected cattle. Results Using serohemorrhagic exudates ( n  = 6), biopsies ( n  = 2) and flies ( n  = 1), the PCR targeting the cox 1 gene resulted in a gel band of almost 700 bp. Cloning, sequencing, and removal of primer sequences yielded a 649-bp fragment of the P. bovicola cox 1 gene. The PCR targeting the ITS region showed a band of about 1100 bp. Cloning, sequencing, and removal of primer sequences resulted in a 1083 bp stretch of the P. bovicola ITS region. Testing samples from presumably affected animals, the cox 1-PCR resulted in bands with the expected size and they were all confirmed as P. bovicola by sequencing. In contrast, the ITS-PCR proved to be less sensitive and less specific and additionally amplified the ITS region of Musca domestica or buttercup DNA. When analysing for sensitivity, the cox 1-PCR yielded visible bands up to 2 ng of genomic DNA, whereas the ITS-PCR produced bands up to 3 ng. In a plasmid dilution series, the minimum number of target DNA copies was 10 2 for the cox 1-PCR and 10 1 in the ITS-PCR. Conclusions The evaluated cox 1-PCR enables reliable detection of P. bovicola DNA in skin biopsies and serohemorrhagic exudates. This PCR and, to a limited extent, the ITS-PCR, may help evaluate different therapeutic approaches. Furthermore, the cox 1-PCR may be useful for epidemiological studies on the geographical distribution of P. bovicola . Further understanding of the epidemiology of this parasite will help develop and implement effective control strategies.

Beneficial effects of monotherapy with ACE inhibitors or beta-blockers on hemodynamic function after myocardial infarction are well known. Until now, the effects of combined treatment on cardiac function and energy metabolism have been poorly described. This study examines the effects of combined ramipril and metoprolol treatment on the creatine kinase (CK) system and hemodynamic function in rats after infarction. Wistar rats with experimental infarction were randomized for treatment with ramipril (R), metoprolol (M), combined treatment (MR), or placebo (P). Sham-operated (SO) animals served as controls. After 6 weeks, we assayed for CK isoenzymes and performed hemodynamic measurements. In P versus SO, left ventricular systolic pressures (dp/dt(max) and dp/dt(min)) diminished, whereas left ventricular end-diastolic pressure (LVEDP) increased. Decreased total CK activity and mitochondrial CK isoenzyme, increased CK-MB, and increased CK-BB isoenzymes were measured in P versus SO. With infarct size < or =45%, mitochondrial CK increased in M and R versus P. Combined treatment had an additional enhancing effect on mitochondrial CK isoenzyme level versus M and R, decreased LVEDP versus P, as well as increased dp/dt(max) and dp/dt(min) versus R. These results provide evidence of an interaction between normalization of energy metabolism and improvement in cardiac function due to a combination of ACE inhibition and beta blockade after myocardial infarction.