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The process of making stone tools, specifically knapping, is a hominin behaviour that typically involves using the upper limb to manipulate a stone hammer and apply concentrated percussive force to another stone, causing fracture and detachment of stone chips with sharp edges. To understand the emergence and subsequent evolution of tool-related behaviours in hominins, the connections between the mechanics of stone knapping, including the delivery of percussive forces, and biomechanics and hominin anatomy, especially in the upper limb, are required. However, there is an absence of direct experimental means to measure the actual forces generated and applied to produce flakes during knapping. Our study introduces a novel solution to this problem in the form of an ergonomic hand-held synthetic hammerstone that can record the percussive forces that occur during knapping experiments. This hammerstone is composed of a deformable pneumatic 3D-printed chamber encased within a 3D-printed grip and a stone-milled striker. During knapping, hammer impact causes the pneumatic chamber to deform, which leads to a change in pressure that is measured by a sensor. Comparisons of recorded pressure data against corresponding force values measured using a force plate show that the synthetic hammer quantifies percussion forces with relatively high accuracy. The performance of this hammerstone was further validated by conducting anvil-supported knapping experiments on glass that resulted in a root mean square error of under 6%, while recording forces up to 730 N with successful flake detachments. These validation results indicate that accuracy was not sensitive to variations up to 15° from the vertical in the hammer striking angle. Our approach allows future studies to directly examine the role of percussive force during the stone knapping process and its relationship with both anatomical and technological changes during human evolution.

New excavations in Liang Bua, where the remains of the ‘Hobbit’ ( Homo floresiensis ) were discovered, show that this diminutive human species used this cave between 190,000 and 50,000 years ago, and not until as recently as 12,000 years ago as previously interpreted; modern humans have been present in Australia since around 50,000 years ago, so whether Homo floresiensis survived long enough to witness the arrival of modern humans is still an open question. An earlier date for Homo floresiensis The discovery in Liang Bua cave on the island of Flores in Indonesia of the diminutive Homo floresiensis , an archaic member of the human family commonly known as the 'Hobbit', was an archaeological sensation in 2004. A source of considerable debate was the fact that it lived in Liang Bua cave between 95,000 and 12,000 years ago, after modern humans had colonized the area (around 50,000 years ago). Thomas Sutikna and colleagues — including many of the original research team — have gone back to Liang Bua, where new excavations have exposed previously unexplored parts of the cave. They have found that the layers of sediment in the cave are not deposited evenly, and it now seems that the H. floresiensis -bearing strata are older than was thought. New radiometric dating places the H. floresiensis remains and stone artefacts to between 190,000 and 50,000 years ago. Whether H. floresiensis survived long enough to witness the arrival of modern humans is an open question. Homo floresiensis , a primitive hominin species discovered in Late Pleistocene sediments at Liang Bua (Flores, Indonesia) 1 , 2 , 3 , has generated wide interest and scientific debate. A major reason this taxon is controversial is because the H. floresiensis -bearing deposits, which include associated stone artefacts 2 , 3 , 4 and remains of other extinct endemic fauna 5 , 6 , were dated to between about 95 and 12 thousand calendar years (kyr) ago 2 , 3 , 7 . These ages suggested that H. floresiensis survived until long after modern humans reached Australia by ~50 kyr ago 8 , 9 , 10 . Here we report new stratigraphic and chronological evidence from Liang Bua that does not support the ages inferred previously for the H. floresiensis holotype (LB1), ~18 thousand calibrated radiocarbon years before present (kyr cal. bp ), or the time of last appearance of this species (about 17 or 13–11 kyr cal. bp ) 1 , 2 , 3 , 7 , 11 . Instead, the skeletal remains of H. floresiensis and the deposits containing them are dated to between about 100 and 60 kyr ago, whereas stone artefacts attributable to this species range from about 190 to 50 kyr in age. Whether H. floresiensis survived after 50 kyr ago—potentially encountering modern humans on Flores or other hominins dispersing through southeast Asia, such as Denisovans 12 , 13 —is an open question.

A nearly complete right hand of an adult hominin was recovered from the Rising Star cave system, South Africa. Based on associated hominin material, the bones of this hand are attributed to Homo naledi . This hand reveals a long, robust thumb and derived wrist morphology that is shared with Neandertals and modern humans, and considered adaptive for intensified manual manipulation. However, the finger bones are longer and more curved than in most australopiths, indicating frequent use of the hand during life for strong grasping during locomotor climbing and suspension. These markedly curved digits in combination with an otherwise human-like wrist and palm indicate a significant degree of climbing, despite the derived nature of many aspects of the hand and other regions of the postcranial skeleton in H. naledi . It is unclear to what extent early hominins were adapted to arboreal climbing. Here, the authors show that the nearly complete hand of H. naledi from South Africa has markedly curved digits and otherwise human-like wrist and palm, which indicates the retention of a significant degree of climbing.

Despite discoveries of relatively complete hands from two early hominin species (Ardipithecus ramidus and Australopithecus sediba) and partial hands from another (Australopithecus afarensis), fundamental questions remain about the evolution of human-like hand anatomy and function. These questions are driven by the paucity of hand fossils in the hominin fossil record between 800,000 and 1.8 My old, a time interval well documented for the emergence and subsequent proliferation of Acheulian technology (shaped bifacial stone tools). Modern and Middle to Late Pleistocene humans share a suite of derived features in the thumb, wrist, and radial carpometacarpal joints that is noticeably absent in early hominins. Here we show that one of the most distinctive features of this suite in the Middle Pleistocene to recent human hand, the third metacarpal styloid process, was present ∼1.42 Mya in an East African hominin from Kaitio, West Turkana, Kenya. This fossil thus provides the earliest unambiguous evidence for the evolution of a key shared derived characteristic of modern human and Neandertal hand morphology and suggests that the distinctive complex of radial carpometacarpal joint features in the human hand arose early in the evolution of the genus Homo and probably in Homo erectus sensu lato .

Compared with H. erectus, these species - for example, other early Homo species such as Homo habilis, as well as the australopiths (hominins not in the genus Homo), which include Paranthropus and Australopithecus - had smaller brains and an anatomy that is less similar to that of modern humans. [...]the H. luzonensis finger and toe bones are curved, which suggests that climbing was an important part of this species' behavioural repertoire, as was also the case for many species of early hominin9. [...]it is worth considering how different these ideas might be if, in the 1890s, H. floresiensis or H. luzonensis had been discovered rather than H. erectus. Because H. luzonensis provides the first glimpse of a second hominin species living on a distant island at a time when H. sapiens populations from Africa were beginning to spread across the world, one thing can be said for certain - our picture of hominin evolution in Asia during the Pleistocene just got even messier, more complicated and a whole lot more interesting. ?

The origin of the human lineage was catalyzed by bipedalism, but how this locomotor mode evolved is debated. We investigated the evolutionary context of human bipedalism by analyzing the morphology of the 4.4 million-year-old hominin talus attributed to Ardipithecus ramidus (ARA-VP-6/500-023). Our results demonstrate that ARA-VP-6/500-023 bears similarities to the tali of chimpanzees and gorillas, who are adapted to vertical climbing and terrestrial plantigrade quadrupedalism. Additionally, we identify the presence of derived features in ARA-VP-6/500-023 consistent with previous suggestions of an enhanced push-off mechanism in the foot of Ar. ramidus . Our observations of the human and ape fossil record are inconsistent with recently proposed models of human origins, which envision the last common ancestor of humans and chimpanzees as a generalized arboreal ape. Instead, our results strongly imply that humans evolved from an African ape-like ancestor, which directly narrows the range of explanations for the origin of our lineage. Morphometric analyses of ankle bones provide evidence that humans evolved from an ancestor with vertical climbing adaptations like those of chimpanzees and gorillas.

Excavations in southeast Asia have unearthed a previously unreported hominin species named Homo luzonensis . The discovery has implications for ideas about early hominin evolution and dispersal from Africa. The discovery of a species of hominin called Homo luzonensis.

The field of phenomics is experiencing unprecedented advances thanks to the rapid growth of morphological quantification based on three-dimensional (3D) imaging, online data repositories, team-oriented collaborations, and open data-sharing policies. In line with these progressions, we present an extensive primate phenotypic dataset comprising >6,000 3D scans (media) representing skeletal morphologies of 386 individual specimens covering all hominoid genera (except humans) and other selected primates. The digitized specimens are housed in physical collections at the American Museum of Natural History, the National Museum of Natural History, the Royal Museum for Central Africa (Belgium), the Cleveland Museum of Natural History, and Stony Brook University. Our technical validation indicates that despite the diverse digitizing devices used to produce the scans, the final 3D models (meshes) can be safely combined to collect comparable morphometric data. The entire dataset (and detailed associated metadata) is freely available through MorphoSource. Hence, these data contribute to empowering the future of primate phenomics and providing a roadmap for future digitization and archiving of digital data from other collections.

Background: Passerines (Aves: Passeriformes) dominate modern terrestrial bird communities yet their fossil record is limited. Liang Bua is a large cave on the Indonesian island of Flores that preserves Late Pleistocene–Holocene deposits (∼190 ka to present day). Birds are the most diverse faunal group at Liang Bua and are present throughout the stratigraphic sequence. Methods: We examined avian remains from the Late Pleistocene deposits of Sector XII, a 2 × 2 m area excavated to about 8.5 m depth. Although postcranial passerine remains are typically challenging to identify, we found several humeral characters particularly useful in discriminating between groups, and identified 89 skeletal elements of passerines. Results: At least eight species from eight families are represented, including the Large-billed Crow (Corvus cf. macrorhynchos), the Australasian Bushlark (Mirafra javanica), a friarbird (Philemon sp.), and the Pechora Pipit (Anthus cf. gustavi). Discussion: These remains constitute the first sample of fossil passerines described in Wallacea. Two of the taxa no longer occur on Flores today; a large sturnid (cf. Acridotheres) and a grassbird (Megalurus sp.). Palaeoecologically, the songbird assemblage suggests open grassland and tall forests, which is consistent with conditions inferred from the non-passerine fauna at the site. Corvus cf. macrorhynchos, found in the Homo floresiensis-bearing layers, was likely part of a scavenging guild that fed on carcasses of Stegodon florensis insularis alongside vultures (Trigonoceps sp.), giant storks (Leptoptilos robustus), komodo dragons (Varanus komodoensis), and probably H. floresiensis as well.

Organic biomarker and lithic use-wear analyses of archaeological implements manufactured and/or used by hominins in the past offers a means of assessing how prehistoric peoples utilised natural resources. Currently, most studies focus on one of these techniques, rather than using both in sequence. This study aims to assess the potential of combining both methods to analyse stone artefacts, using a set of 69 stones excavated from the cave site of Liang Bua (Flores, Indonesia). Prior to chemical analysis, an initial inspection of the artefacts revealed potential use-wear traces but no visible residues. Gas chromatography mass spectrometry (GC-MS) analysis, including the targeting of 86 lipids, terpenes, terpenoids, alkanes and their analogues, found compounds with plant or animal origin on 27 of the 69 stones. The artefacts were subsequently cleaned, and use-wear analysis identified traces of use on 43 artefacts. Use-wear analysis confirmed traces of use on 23 of the 27 artefacts with potential use-residues that were determined by GC-MS. The GC-MS results were broadly consistent with the functional classes identified in the later use-wear analysis. This inclusive approach for stone artefact analysis strengthens the identifications made through multiple lines of enquiry. There remain conflicts and uncertainties in specific cases, suggesting the need for further refinement and analyses of the relationships between use-wear and residues.