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Euchelicerata is a clade of arthropods comprising horseshoe crabs, scorpions, spiders, mites and ticks, as well as the extinct eurypterids (sea scorpions) and chasmataspidids. The understanding of the ground plans and relationships between these crown-group euchelicerates has benefited from the discovery of numerous fossils. However, little is known regarding the origin and early evolution of the euchelicerate body plan because the relationships between their Cambrian sister taxa and synziphosurines, a group of Silurian to Carboniferous stem euchelicerates with chelicerae and an unfused opisthosoma, remain poorly understood owing to the scarce fossil record of appendages. Here we describe a synziphosurine from the Lower Ordovician (ca. 478 Ma) Fezouata Shale of Morocco. This species possesses five biramous appendages with stenopodous exopods bearing setae in the prosoma and a fully expressed first tergite in the opisthosoma illuminating the ancestral anatomy of the group. Phylogenetic analyses recover this fossil as a member of the stem euchelicerate family Offacolidae, which is characterized by biramous prosomal appendages. Moreover, it also shares anatomical features with the Cambrian euarthropod Habelia optata , filling the anatomical gap between euchelicerates and Cambrian stem taxa, while also contributing to our understanding of the evolution of euchelicerate uniramous prosomal appendages and tagmosis. Here, the authors describe an early synziphosurine from the Lower Ordovician Fezouata Shale of Morocco, which exhibits traits that elucidate the long-contentious relationships between crown euchelicerates and their sister taxa, and also clarifies euchelicerate body plan evolution.

Spatial learning is a complex cognitive skill and ecologically important trait scarcely studied in crustaceans. We investigated the ability of the Pacific (Ecuadorian) hermit crab Coenobita compressus, to learn an allocentric spatial task using a palatable novel food as reward. Crabs were trained to locate the reward in a single session of eleven consecutive trials and tested subsequently, for short- (5 min) and long-term memory 1, 3 and 7 days later. Our results indicate that crabs were able to learn the location of the reward as they showed a reduction in the time required to find the food whenever it was present, suggesting a visuo-spatial and olfactory cue-guided task resolution. Moreover, crabs also remember the location of the reward up to 7 days after training using spatial cues only (without the food), as evidenced by the longer investigation time they spent in the learned food location than in any other part of the experimental arena, suggesting a visuo-spatial memory formation. This study represents the first description of allocentric spatial long-term memory in a terrestrial hermit crab.

Specimens of Euproops sp. (Xiphosura, Chelicerata) from the Carboniferous Piesberg quarry near Osnabrück, Germany, represent a relatively complete growth series of 10 stages. Based on this growth sequence, morphological changes throughout the ontogeny can be identified. The major change affects the shape of the epimera of the opisthosoma. In earlier stages, they appear very spine-like, whereas in later stages the bases of these spine-like structures become broader; the broadened bases are then successively drawn out distally. In the most mature stage known, the epimera are of trapezoidal shape and approach each other closely to form a complete flange around the thoracetron (=fused tergites of the opisthosoma). These ontogenetic changes question the taxonomic status of different species of Euproops, as the latter appear to correspond to different stages of the ontogenetic series reconstructed from the Piesberg specimens. This means that supposed separate species could, in fact, represent different growth stages of a single species. It could alternatively indicate that heterochrony (=evolutionary change of developmental timing) plays an important role in the evolution of Xiphosura. We propose a holomorph approach, i.e., reconstructing ontogenetic sequences for fossil and extant species as a sound basis for a taxonomic, phylogenetic, and evolutionary discussion of Xiphosura.

The world's nearly 7,000 species of crabs are immediately recognizable by their claws, sideways movement, stalked eyes, and thick outer shells. These common crustaceans are found internationally, thriving in various habitats from the edge of the sea to the depths of the ocean, in fresh water or on land. Despite having the same basic body type as decapod crustaceans-true crabs have heavy exoskeletons and ten limbs with front pincer claws-crabs come in an enormous variety of shapes and sizes, from the near microscopic to the giant Japanese spider crab. InWalking Sideways, Judith S. Weis provides an engaging and informative tour of the remarkable world of crabs, highlighting their unique biology and natural history. She introduces us to recently discovered crabs such as the Yeti crab found in deep sea vents, explains what scientists are learning about blue and hermit crabs commonly found at the shore, and gives us insight into the lifecycles of the king and Dungeness crabs typically seen only on dinner plates. Among the topics Weis covers are the evolution and classification of crabs, their habitats, unique adaptations to water and land, reproduction and development, behavior, ecology, and threats, including up-to-date research. Crabs are of special interest to biologists for their communication behaviors, sexual dimorphism, and use of chemical stimuli and touch receptors, and Weis explains the importance of new scientific discoveries. In addition to the traditional ten-legged crabs, the book also treats those that appear eight-legged, including hermit crabs, king crabs, and sand crabs. Sidebars address topics of special interest, such as the relationship of lobsters to crabs and medical uses of compounds derived from horseshoe crabs (which aren't really crabs). While Weis emphasizes conservation and the threats that crabs face, she also addresses the use of crabs as food (detailing how crabs are caught and cooked) and their commercial value from fisheries and aquaculture. She highlights other interactions between crabs and people, including keeping hermit crabs as pets or studying marine species in the laboratory and field. Reminding us of characters such asThe Little Mermaid's Sebastian and Sherman Lagoon's Hawthorne, she also surveys the role of crabs in literature (for both children and adults), film, and television, as well in mythology and astrology. With illustrations that offer delightful visual evidence of crab diversity and their unique behaviors,Walking Sidewayswill appeal to anyone who has encountered these fascinating animals on the beach, at an aquarium, or in the kitchen.

A young and energetic pulsar powers the well-known Crab Nebula. Here, we describe two separate gamma-ray (photon energy greater than 100 mega-electron volts) flares from this source detected by the Large Area Telescope on board the Fermi Gamma-ray Space Telescope. The first flare occurred in February 2009 and lasted approximately 16 days. The second flare was detected in September 2010 and lasted approximately 4 days. During these outbursts, the gamma-ray flux from the nebula increased by factors of four and six, respectively. The brevity of the flares implies that the gamma rays were emitted via synchrotron radiation from peta-electron-volt (10¹⁵ electron volts) electrons in a region smaller than 1.4 x 10⁻² parsecs. These are the highest-energy particles that can be associated with a discrete astronomical source, and they pose challenges to particle acceleration theory.