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1. Priority question exercises are becoming an increasingly common tool to frame future agendas in conservation and ecological science. They are an effective way to identify research foci that advance the field and that also have high policy and conservation relevance. 2. To date, there has been no coherent synthesis of key questions and priority research areas for palaeoecology, which combines biological, geochemical and molecular techniques in order to reconstruct past ecological and environmental systems on time-scales from decades to millions of years. 3. We adapted a well-established methodology to identify 50 priority research questions in palaeoecology. Using a set of criteria designed to identify realistic and achievable research goals, we selected questions from a pool submitted by the international palaeoecology research community and relevant policy practitioners. 4. The integration of online participation, both before and during the workshop, increased international engagement in question selection. 5. The questions selected are structured around six themes: human–environment interactions in the Anthropocene; biodiversity, conservation and novel ecosystems; biodiversity over long time-scales; ecosystem processes and biogeochemical cycling; comparing, combining and synthesizing information from multiple records; and new developments in palaeoecology. 6. Future opportunities in palaeoecology are related to improved incorporation of uncertainty into reconstructions, an enhanced understanding of ecological and evolutionary dynamics and processes and the continued application of long-term data for better-informed landscape management. 7. Synthesis. Palaeoecology is a vibrant and thriving discipline, and these 50 priority questions highlight its potential for addressing both pure (e.g. ecological and evolutionary, methodological) and applied (e.g. environmental and conservation) issues related to ecological science and global change.

As the Earth's Third Pole and the Asian water tower, the Qinghai‐Tibet Plateau (QTP) plays a key role in global climate regulation and biodiversity maintenance. Living in harmony with nature is vital for local and global sustainable development. Current research on the conflicted or coordinated relationship between humans and nature on the QTP at a fine spatial scale remains limited. To fill the gap, we developed the human activity intensity index (HAI) and eco‐environmental quality index (EQI) at 1‐km resolution and proposed a four‐quadrant diagram approach to explore the dynamics between them. The results show a coordinated development on the QTP as the HAI and EQI both increased from 2000 to 2020, and the ratio of coordinated areas to conflicted areas was 5:1. High HAI areas were mainly in big cities such as Xining, Lhasa, Haidong, Xigaze, and along traffic lines. The significant conflicted areas were mainly outside the Lhasa metropolitan, south of the Hengduan Mountains, and along some new roads, and reduced by 8% between 2000–2010 and 2010–2020. The area of high HAI but low EQI was the smallest proportion, mainly in southern Qinghai Lake, southern Brahlung Zangbo River, Gobi oases, and western transport lines, but it implies the highest risk of ecosystem degradation. This research expands the fundamental methodology to address complex human‐natural relationships and provides implications for the sustainable development of fragile ecosystems. Plain Language Summary The Qinghai‐Tibet Plateau (QTP), with an average altitude of over 4,000 m and 13 million residents, is the source of the nine rivers in Asia, providing fresh water, food, and other ecosystem services to more than 1.5 billion people, and is known as the Earth's Third Pole and Asian water tower. However, research on the relationship between humans and nature in that region is limited, especially at a fine spatial scale. To fill the gap, we developed the human activity intensity index (HAI) and eco‐environmental quality index (EQI) at 1‐km resolution and proposed a four‐quadrant diagram approach to explore the dynamics between them, addressing potential risks and sustainability pathways. We find that the relationship between humans and nature on the QTP tends to be harmonious from 2000 to 2020. The significant conflicted areas were mainly outside the Lhasa metropolitan, south of the Hengduan Mountains, and along new roads, and reduced by 8% between 2000–2010 and 2010–2020. However, the plateau's fragile ecosystem still faces great challenges with population growth, urbanization, infrastructure construction, and the threat of global climate change. This work expands the fundamental methodology and may support fine ecological restoration and environmental management for local governments. Key Points Human activity intensity and eco‐environmental quality were measured at the grid scale of 1‐km resolution on the Earth's Third Pole We proposed a four‐quadrant diagram approach to identify dynamic relationships between humans and nature Socioeconomic development and eco‐environment on the QTP tend to be coordinated during 2000–2020

Coral reef fisheries support the livelihoods of millions of people but have been severely and negatively affected by anthropogenic activities. We conducted a systematic review of published data on the biomass of coral reef fishes to explore how the condition of reef fisheries is related to the density of local human populations, proximity of the reef to markets, and key environmental variables (including broad geomorphologic reef type, reef area, and net productivity). When only population density and environmental covariates were considered, high variability in fisheries conditions at low human population densities resulted in relatively weak explanatory models. The presence or absence of human settlements, habitat type, and distance to fish markets provided a much stronger explanatory model for the condition of reef fisheries. Fish biomass remained relatively low within 14 km of markets, then biomass increased exponentially as distance from reefs to markets increased. Our results suggest the need for an increased science and policy focus on markets as both a key driver of the condition of reef fisheries and a potential source of solutions. Las pesquerías en arrecifes de coral soportan a millones de personas pero han sido severa y negativamente afectadas por actividades antropogénicas. Realizamos una revisión sistemática de datos publicados sobre la biomasa de peces de arrecifes de coral para explorar la relación entre la condición de las pesquerías en arrecifes y la densidad de las poblaciones humanas locales, la cercanía del arrecife a los mercados y variables ambientales claves (incluyendo el tipo de arrecife geomorfológico, la superficie del arrecife y la productividad neta). Cuando solo se consideraron covariables de densidad y hábitat, la alta variabilidad en las condiciones de la pesquería a densidades bajas de la población humana resultó en modelos explicativos relativamente débiles. La presencia o ausencia de asentamientos humanos, el tipo de hábitat y la distancia a los mercados de pescado proporcionaron un modelo explicativo mucho más robusto para a condición de las pesquerías en arrecifes. La biomasa de peces permaneció relativamente baja a menos de 14 km de los mercados, pero más allá de 14 km la biomasa incrementó exponencialmente a medida que incrementó la distancia entre arrecifes y mercados. Nuestros resultados sugieren que los mercados, como un factor clave de la condición de las pesquerías en arrecifes y como una potencial fuente de soluciones, requieren una mayor atención de la ciencia y la política.

Worldwide urbanization leads to ecological changes around urban areas. However, few studies have quantitatively investigated the impacts of urbanization on vegetation coverage so far. As an important indicator measuring regional environment change, fractional vegetation cover (FVC) is widely used to analyze changes in vegetation in urban areas. In this study, on the basis of a partial derivative model, we quantified the effect of temperature, precipitation, radiation, and urbanization represented as nighttime light on vegetation coverage changes in the Beijing–Tianjin–Hebei (BTH) region during its period of rapid resident population growth from 2001 to 2011. The results showed that (1) the FVC of the BTH region varied from 0.20 to 0.26, with significant spatial heterogeneity. The FVC increased in small cities such as Cangzhou and in the Taihang Mountains, while it decreased in megacities with populations greater than 1 million, such as Beijing and Zhangjiakou Bashang. (2) The BTH region experienced rapid urbanization, with the area of artificial surface increasing by 18.42%. From the urban core area to the fringe area, the urbanization intensity decreased, but the urbanization rate increased. (3) Urbanization and precipitation had the greatest effect on FVC changes. Urbanization dominated the FVC changes in the expanded area, while precipitation had the greatest impacts on the FVC changes in the core area. For future studies on the major influencing factors of FVC changes, quantitative analysis of the contribution of urbanization to FVC changes in urban regions is crucial and will provide scientific perspectives for sustainable urban planning.

Navigating social‐ecological systems toward sustainable trajectories is an important challenge of the Anthropocene. Models of social‐ecological systems can increase our understanding of how social and ecological subsystems interact, their response to environmental changes, and how their dynamics may be altered by management interventions. However, the level of representational detail required for models to describe a particular social‐ecological system with high fidelity (i.e., accurately quantifying system dynamics) may hamper both the interpretability of model results and our ability to identify key processes and feedbacks within the system. In contrast, stylized models describe simplified interactions between a small subset of social‐ecological system elements. Stylized models are a useful tool to identify potential consequences of specific key processes and feedbacks on system functioning. However, the relatively low level of representational detail in these models limits their ability to deliver concrete management options for a particular social‐ecological system. Here, we describe how an exploratory modeling approach can utilize the strengths of stylized models before the construction of social‐ecological system models with high fidelity and representational detail. This exploratory modeling approach is an iterative strategy, with the initial steps comprising the development of stylized models informed by empirical observations. We illustrate this with two examples of stylized modeling of isolated and connected social‐ecological systems. Through repeated confrontation of alternative models with empirical data, exploratory modeling provides useful stepping stones toward the development of models that describe social‐ecological systems in increasingly specific settings with increasing levels of representational detail. When these latter types of models reach a high level of fidelity, they could be used for scenario‐based analyses and participatory decision‐making processes. At this stage, the conceptual insights previously obtained during the exploratory modeling phase may aid in the interpretation and communication of the outcomes of scenario‐based analyses. Hence, exploratory modeling aims to create a synergy between the insights obtained from stylized models and system‐specific, high‐fidelity models in order to generate a deep understanding of the drivers of social‐ecological system dynamics, and how to leverage these drivers to initiate desired changes.

Population concentration and built‐up land expansion are two prominent features of contemporary urbanization. Existing literature on the population aspect of urbanization has mostly focused on national and regional aggregates, and literature on the land development aspect has often relied on spatial case studies of individual cities or their meta‐analyses. Using newly‐available data, here we conduct the first global‐coverage, spatial analysis of the relationship between (changes in) population and built‐up land at multiple spatial scales, and compare to existing common beliefs about urbanization based on individual city studies. We find that population and built‐up land show distinctly different spatial and temporal patterns (with a global correlation coefficient around 0.6). Contrary to common impressions, our results show that during recent decades, developed and developing regions across the world experienced comparable amounts of built‐up land expansion. While meta‐analyses have reported that built‐up land in urban areas expands globally on average twice as fast as population grows, our results show the global change rates of built‐up land and population are similar. Also, most global population, including what national statistics agencies call urban population, reside in areas with low land development levels (which are frequently less than 5% built up). These changes in perspective suggest that urbanization's potential large‐scale impacts may need to be re‐evaluated, and lead to best‐practice recommendations for urbanization modeling and analysis. Especially, the common practice in large‐scale earth system modeling of assuming demographically‐defined urban population resides in areas with medium to high built‐up land development levels should change. Plain Language Summary Cities and towns across the world have been expanding (a process known as urbanization) both in terms of population size and land areas. These two expressions of urbanization (i.e., population concentration and built‐up land expansion) have often been assumed to be strongly associated. This assumption is influential for understanding people's vulnerability to natural disasters (for example, heatwaves, coastal storms) in urban areas, but has not been tested globally, largely due to the lack of data. Recently new global‐coverage maps of population and built‐up land became available. Using the newly‐available data, we examine the relationship between population and built‐up land across the world. We find the correlation between population and built‐up land is about 60%, with the two showing some distinctly different spatial and temporal patterns. Contrary to common impressions, we find developed and developing regions across the world experienced similar amounts of built‐up land expansion during recent decades. Moreover, most people in the world (including those considered urban people by national agencies, such as census) live in areas with less than 5% built‐up land. These changes in perspective suggest that urbanization's potential large‐scale impacts may need to be re‐evaluated. Key Points The developed and the developing world experienced comparable amounts of built‐up land expansion during recent decades Most global population (including urban population) reside in areas with low land development levels (frequently less than 5% built up) Population and built‐up land show distinctly different spatiotemporal patterns globally (with a correlation coefficient around 0.6)

AIM: Biologists increasingly recognize the roles of humans in ecosystems. Subsequently, many have argued that biodiversity conservation must be extended to environments that humans have shaped directly. Yet popular biogeographical frameworks such as biomes do not incorporate human land use, limiting their relevance to future conservation planning. ‘Anthromes’ map global ecological patterns created by sustained direct human interactions with ecosystems. In this paper, we set to understand how current conservation efforts are distributed across anthromes. LOCATION: Global. METHODS: We analysed the global distribution of IUCN protected areas and biodiversity hotspots by anthrome. We related this information to density of native plant species and density of previous ecological studies. Potential conservation opportunities in anthromes were then identified through global analysis and two case studies. RESULTS: Protected areas and biodiversity hotspots are not distributed equally across anthromes. Less populated anthromes contain a greater proportion of protected areas. The fewest hotspots are found within densely settled anthromes and wildlands, which occur at the two extremes of human population density. Opportunities for representative protection, prioritization, study and inclusion of native species were not congruent. MAIN CONCLUSIONS: Researchers and practitioners can use the anthromes framework to analyse the distribution of conservation practices at the global and regional scale. Like biomes, anthromes could also be used to set future conservation priorities. Conservation goals in areas directly shaped by humans need not be less ambitious than those in ‘natural areas’.

The monumental Late Bonze Age royal tomb of Seddin is located in the old morainic landscape of the Prignitz region, northeastern Germany. Together with other richly equipped burials and a row of stone pits in its direct vicinity, it provides evidence for the presence of an elite from the nineth to sixth centuries BCE in this region. Our map emphasizes the well-chosen location of the royal tomb in relation to the spatial arrangement of other archaeological monuments that together form an ensemble of a ritual landscape. We trace legacies of land use from the Bronze Age to the present against the backdrop of Late Quaternary landscape evolution. These include the Bronze Age landscape (re-)organization for ritual and economic purposes, its medieval use for arable farming, its economic use and settlement history in historic times, and modern times melioration of agricultural areas that together form the palimpsest of the present-day landscape.