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Recent climate changes have increased fire-prone weather conditions in many regions and have likely affected fire occurrence, which might impact ecosystem functioning, biogeochemical cycles, and society. Prediction of how fire impacts may change in the future is difficult because of the complexity of the controls on fire occurrence and burned area. Here we aim to assess how process-based fire-enabled dynamic global vegetation models (DGVMs) represent relationships between controlling factors and burned area. We developed a pattern-oriented model evaluation approach using the random forest (RF) algorithm to identify emergent relationships between climate, vegetation, and socio-economic predictor variables and burned area. We applied this approach to monthly burned area time series for the period from 2005 to 2011 from satellite observations and from DGVMs from the “Fire Modeling Intercomparison Project” (FireMIP) that were run using a common protocol and forcing data sets. The satellite-derived relationships indicate strong sensitivity to climate variables (e.g. maximum temperature, number of wet days), vegetation properties (e.g. vegetation type, previous-season plant productivity and leaf area, woody litter), and to socio-economic variables (e.g. human population density). DGVMs broadly reproduce the relationships with climate variables and, for some models, with population density. Interestingly, satellite-derived responses show a strong increase in burned area with an increase in previous-season leaf area index and plant productivity in most fire-prone ecosystems, which was largely underestimated by most DGVMs. Hence, our pattern-oriented model evaluation approach allowed us to diagnose that vegetation effects on fire are a main deficiency regarding fire-enabled dynamic global vegetation models' ability to accurately simulate the role of fire under global environmental change.

Several studies have shown that hydrological models do not perform well when applied to periods with climate conditions that differ from those during model calibration. This has important implications for the application of these models in climate change impact studies. The causes of the low transferability to changed climate conditions have, however, only been investigated in a few studies. Here we revisit a study in Austria that demonstrated the inability of a conceptual semi-distributed HBV-type model to simulate the observed discharge response to increases in precipitation and air temperature. The aim of the paper is to shed light on the reasons for these model problems. We set up hypotheses for the possible causes of the mismatch between the observed and simulated changes in discharge and evaluate these using simulations with modifications of the model. In the baseline model, trends of simulated and observed discharge over 1978–2013 differ, on average over all 156 catchments, by 95±50 mm yr−1 per 35 years. Accounting for variations in vegetation dynamics, as derived from a satellite-based vegetation index, in the calculation of reference evaporation explains 36±9 mm yr−1 per 35 years of the differences between the trends in simulated and observed discharge. Inhomogeneities in the precipitation data, caused by a variable number of stations, explain 39±26 mm yr−1 per 35 years of this difference. Extending the calibration period from 5 to 25 years, including annually aggregated discharge data or snow cover data in the objective function, or estimating evaporation with the Penman–Monteith instead of the Blaney–Criddle approach has little influence on the simulated discharge trends (5 mm yr−1 per 35 years or less). The precipitation data problem highlights the importance of using precipitation data based on a stationary input station network when studying hydrologic changes. The model structure problem with respect to vegetation dynamics is likely relevant for a wide spectrum of regions in a transient climate and has important implications for climate change impact studies.

Volatile organic compounds (VOCs) are released from biogenic sources in a temperature-dependent manner. Consequently, Arctic ecosystems are expected to greatly increase their VOC emissions with ongoing climate warming, which is proceeding at twice the rate of global temperature rise. Here, we show that ongoing warming has strong, increasing effects on Arctic VOC emissions. Using a combination of statistical modeling on data from several warming experiments in the Arctic tundra and dynamic ecosystem modeling, we separate the impacts of temperature and soil moisture into direct effects and indirect effects through vegetation composition and biomass alterations. The indirect effects of warming on VOC emissions were significant but smaller than the direct effects, during the 14-y model simulation period. Furthermore, vegetation changes also cause shifts in the chemical speciation of emissions. Both direct and indirect effects result in large geographic differences in VOC emission responses in the warming Arctic, depending on the local vegetation cover and the climate dynamics. Our results outline complex links between local climate, vegetation, and ecosystem–atmosphere interactions, with likely local-to-regional impacts on the atmospheric composition.

In recent years, multiple disturbances have significantly altered terrestrial ecosystems in arid and semi-arid regions, particularly on the Mongolian Plateau (MP). Net primary productivity (NPP) of vegetation is an essential component of the surface carbon cycle. As such, it characterizes the state of variation in terrestrial ecosystems and reflects the productive capacity of natural vegetation. This study revealed the complex relationship between the natural environment and NPP in the ecologically fragile and sensitive MP. The modified Carnegie–Ames–Stanford Approach (CASA) model was used to simulate vegetation NPP. Further, the contributions of topography, vegetation, soils, and climate to NPP’s distribution and spatiotemporal variation were explored using the geographic detector model (GDM) and structural equation model (SEM). The study’s findings indicate the following: (1) NPPs for different vegetation types in the MP were in the order of broad-leaved forest > meadow steppe > coniferous forest > cropland > shrub > typical steppe > sandy land > alpine steppe > desert steppe. (2) NPP showed an increasing trend during the growing seasons from 2000 to 2019, with forests providing larger vegetation carbon stocks. It also maintained a more stable level of productivity. (3) Vegetation cover, precipitation, soil moisture, and solar radiation were the key factors affecting NPP’s spatial distribution. NPP’s spatial distribution was primarily explained by the normalized difference vegetation index, solar radiation, precipitation, vegetation type, soil moisture, and soil type (-statistics = 0.86, 0.71, 0.67, 0.67, 0.57, and 0.57, respectively); the contribution of temperature was small (-statistics = 0.26), and topographic factors had the least influence on NPP’s distribution, as their contribution amounted to less than 0.20. (4) A SEM constructed based on the normalized difference vegetation index (NDVI), solar radiation, precipitation, temperature, and soil moisture explained 17% to 65% of the MP’s NPP variations. The total effects of the MP’s NPP variations in absolute values were in the order of NDVI (0.47) > precipitation (0.33) > soil moisture (0.16) > temperature (0.14) > solar radiation (0.02), and the mechanisms responsible for NPP variations differed slightly among the relevant vegetation types. Overall, this study can help understand the mechanisms responsible for the MP’s NPP variations and offer a new perspective for regional vegetation ecosystem management.

Seasonal climate predictions of spring (March‒April‒May) dust weather frequency (DWF) over North China (DWFNC) are conducted based on a previous-summer (June–July–August) normalized difference vegetation index in North China (NDVINC), winter (December–January–February) sea-ice cover index over the Barents Sea (SICBS), and winter Antarctic Oscillation index (AAOI). The year-to-year increment approach is applied to improve the prediction skill. Two statistical prediction schemes—statistical models based on year-to-year-increment-form predictors (SM-DY) and anomaly-form predictors (SM-A)—are applied based on NDVINC, SICBS, and AAOI. The results show that the prediction model using the year-to-year increment approach performs much better in predicting DWFNC, with the correlation coefficient between the average DWFNC and the cross-validated results of SM-DY (SM-A) being 0.80 (0.68) during 1983–2016. A hybrid dynamical–statistical prediction model (HM-DY) is constructed based on NDVINC, SICBS, and a spring 850-hPa geopotential height index, derived from the second version of the NCEP Climate Forecast System. Results show that HM-DY has comparable prediction skill with SM-DY. Both SM-DY and HM-DY are extended to hindcast DWF over the 245 stations in the whole of northern China, indicating comparably high skill. The results show that NDVINC and SICBS account for large variances of the dust climate over northern China. In particular, NDVINC and SICBS can enhance 64% of stations in North China in their prediction of dust climate.

Models of plant water fluxes have evolved from studies focussed on understanding the detailed structure and functioning of specific components of the soil–plant–atmosphere (SPA) continuum to architectures often incorporated inside eco-hydrological and terrestrial biosphere (TB) model schemes. We review here the historical evolution of this field, examine the basic structure of a simplified individual-based model of plant water transport, highlight selected applications for specific ecological problems and conclude by examining outstanding issues requiring further improvements in modelling vegetation water fluxes. We particularly emphasise issues related to the scaling from tissue-level traits to individual-based predictions of water transport, the representation of nonlinear and hysteretic behaviour in soil–xylem hydraulics and the need to incorporate knowledge of hydraulics within broader frameworks of plant ecological strategies and their consequences for predicting community demography and dynamics.

The authors describe carbon system formulation and simulation characteristics of two new global coupled carbon–climate Earth System Models (ESM), ESM2M and ESM2G. These models demonstrate good climate fidelity as described in part I of this study while incorporating explicit and consistent carbon dynamics. The two models differ almost exclusively in the physical ocean component; ESM2M uses the Modular Ocean Model version 4.1 with vertical pressure layers, whereas ESM2G uses generalized ocean layer dynamics with a bulk mixed layer and interior isopycnal layers. On land, both ESMs include a revised land model to simulate competitive vegetation distributions and functioning, including carbon cycling among vegetation, soil, and atmosphere. In the ocean, both models include new biogeochemical algorithms including phytoplankton functional group dynamics with flexible stoichiometry. Preindustrial simulations are spun up to give stable, realistic carbon cycle means and variability. Significant differences in simulation characteristics of these two models are described. Because of differences in oceanic ventilation rates, ESM2M has a stronger biological carbon pump but weaker northward implied atmospheric CO₂ transport than ESM2G. The major advantages of ESM2G over ESM2M are improved representation of surface chlorophyll in the Atlantic and Indian Oceans and thermocline nutrients and oxygen in the North Pacific. Improved tree mortality parameters in ESM2G produced more realistic carbon accumulation in vegetation pools. The major advantages of ESM2M over ESM2G are reduced nutrient and oxygen biases in the southern and tropical oceans.

• Land surface models (LSMs) typically use empirical functions to represent vegetation responses to soil drought. These functions largely neglect recent advances in plant ecophysiology that link xylem hydraulic functioning with stomatal responses to climate. • We developed an analytical stomatal optimization model based on xylem hydraulics (SOX) to predict plant responses to drought. Coupling SOX to the Joint UK Land Environment Simulator (JULES) LSM, we conducted a global evaluation of SOX against leaf- and ecosystem-level observations. • SOX simulates leaf stomatal conductance responses to climate for woody plants more accurately and parsimoniously than the existing JULES stomatal conductance model. An ecosystem-level evaluation at 70 eddy flux sites shows that SOX decreases the sensitivity of gross primary productivity (GPP) to soil moisture, which improves the model agreement with observations and increases the predicted annual GPP by 30% in relation to JULES. SOX decreases JULES root-mean-square error in GPP by up to 45% in evergreen tropical forests, and can simulate realistic patterns of canopy water potential and soil water dynamics at the studied sites. • SOX provides a parsimonious way to incorporate recent advances in plant hydraulics and optimality theory into LSMs, and an alternative to empirical stress factors.