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Grasslands are among the planet’s most imperilled ecosystems, largely because habitat conversion has caused extreme biodiversity loss. In response, managers and scientists aim to recreate grassland habitat, yet these reconstructed grasslands are often species‐poor and lose diversity through time. One potential mechanism to promote biodiversity in grasslands is spillover, or the targeted dispersal of species across habitat boundaries from areas of high to low biodiversity. There is potential for native species to disperse via spillover from high‐quality remnant habitat and establish in reconstructions, thus increasing biodiversity. However, plant dispersal and establishment are often context dependent, and the conditions that promote spillover in grasslands are largely unknown. Here we examine the contexts under which spillover can enhance biodiversity in grasslands. Specifically, we investigate whether the species richness of reconstructions and individual plant dispersal traits alter spillover. To do so, we surveyed plant species richness at reconstructed grasslands of varying diversity adjacent to remnant grasslands. We found that spillover from remnants supplies reconstructions with rare species that would otherwise not be present, but only in reconstructions with lower overall richness. Furthermore, spillover was more likely to occur for species with wind dispersed seeds than species with unassisted seed dispersal. Synthesis and applications. Our results show that the context dependency of both dispersal and establishment processes are critical to understanding when and where spillover can promote biodiversity in reconstructed systems. Understanding these contexts will help land managers leverage natural dispersal to mitigate biodiversity loss by anticipating which species are likely to arrive in reconstructions without assistance and when they are likely to establish. Our results show that the context dependency of both dispersal and establishment processes are critical to understanding when and where spillover can promote biodiversity in reconstructed systems. Understanding these contexts will help land managers leverage natural dispersal to mitigate biodiversity loss by anticipating which species are likely to arrive in reconstructions without assistance and when they are likely to establish.

Ammonites are among the best-known fossils of the Phanerozoic, yet their habitat is poorly understood. Three common ammonite families (Baculitidae, Scaphitidae, and Sphenodiscidae) co-occur with well-preserved planktonic and benthic organisms at the type locality of the upper Maastrichtian Owl Creek Formation, offering an excellent opportunity to constrain their depth habitats through isotopic comparisons among taxa. Based on sedimentary evidence and the micro- and macrofauna at this site, we infer that the 9-m-thick sequence was deposited at a paleodepth of 70–150 m. Taxa present throughout the sequence include a diverse assemblage of ammonites, bivalves, and gastropods, abundant benthic foraminifera, and rare planktonic foraminifera. No stratigraphic trends are observed in the isotopic data of any taxon, and thus all of the data from each taxon are considered as replicates. Oxygen isotope-based temperature estimates from the baculites and scaphites overlap with those of the benthos and are distinct from those of the plankton. In contrast, sphenodiscid temperature estimates span a range that includes estimates of the planktonic foraminifera and of the warmer half of the benthic values. These results suggest baculites and scaphites lived close to the seafloor, whereas sphenodiscids sometimes inhabited the upper water column and/or lived closer to shore. In fact, the rarity and poorer preservation of the sphenodiscids relative to the baculites and scaphites suggests that the sphenodiscid shells may have only reached the Owl Creek locality by drifting seaward after death.

Aim The biogeographical and habitat history of the species‐rich angiosperm genus Artabotrys is reconstructed to assess hypotheses relating to processes that underlie palaeotropical intercontinental disjunction (PID) and regional diversification patterns. Location Palaeotropics. Taxon Artabotrys (Annonaceae). Methods Phylogenetic relationships were estimated based on 53 Artabotrys species, using four chloroplast and 10 nuclear markers (c. 15.7 kb). Divergence times were estimated using two fossil calibrations and an uncorrelated lognormal relaxed clock model. Ancestral range estimation was performed under a dispersal–extinction–cladogenesis model while ancestral habitat reconstruction was conducted using the BAYAREALIKE model. Results Artabotrys is unequivocally monophyletic, with a species‐rich main Artabotrys clade (MAC) comprising distinct African and Asian sister clades, and an early divergent grade (EDG) comprising two African species. An ancestral range in Africa is inferred, with a single dispersal to Asia. The PID at the MAC crown occurred in the Miocene. A broad habitat tolerance spanning rain forests and seasonally dry forests/savannas was inferred at the MAC stem and crown nodes. Several shifts from rain forests to seasonally dry habitats were inferred, but there is no indication of a reverse transition. Main conclusions The most plausible explanation for the PID involves overland migration across Arabia in the Miocene, prior to subsequent climate deterioration. Long‐standing differences in climatic niche may have resulted in a significant yet porous biogeographical divide at the Isthmus of Kra, but Wallace's line does not reflect differences in climatic niches. Niche conservatism is an underlying pattern in Artabotrys, with local niche shifts occurring rather recently.

Late Neogene aridification in the Southern Hemisphere caused contractions of mesic biota to refugia, similar to the patterns established by glaciation in the Northern Hemisphere, but these episodes also opened up new adaptive zones that spurred range expansion and diversification in arid-adapted lineages. To understand these dynamics, we present a multilocus (nine nuclear ¡ ntrons, one mitochondrial gene) phylogeographic analysis of the Bynoe's gecko (Heteronotia binoei), a widely distributed complex spanning the tropical monsoon, coastal woodland, and arid zone biomes in Australia. Bayesian phylogenetic analyses, estimates of divergence times, and demographic inferences revealed episodes of diversification in the Pliocene, especially in the tropical monsoon biome, and range expansions in the Pleistocene. Ancestral habitat reconstructions strongly support recent and independent invasions into the arid zone. Our study demonstrates the varied responses to aridification in Australia, including localized persistence of lineages in the tropical monsoonal biome, and repeated invasion of and expansion through newly available aridzone habitats. These patterns are consistent with those found in other arid environments in the Southern Hemisphere, including the South African succulent karoo and the Chilean lowlands, and highlight the diverse modes of diversification and persistence of Earth's biota during the glacial cycles of the Pliocene and Pleistocene.

Modern flammable ecosystems include tropical and subtropical savannas, steppe grasslands, boreal forests, and temperate sclerophyll shrublands. Despite the apparent fiery nature of much contemporary vegetation, terrestrial fossil evidence would suggest we live in a time of low fire activity relative to the deep past. The inertinite content of coal, fossil charcoal, is strikingly low from the Eocene to the Pleistocene and no charcoalified mesofossils have been reported for the Cenozoic. Marine cores have been analyzed for charcoal in the North Pacific, the north and south Atlantic off Africa, and the south China sea. These tell a different story with the oldest records indicating low levels of fire activity from the Eocene but a surge of fire from the late Miocene (~7 Ma). Phylogenetic studies of woody plants adapted to frequent savanna fires show them beginning to appear from the Late Miocene with peak origins in the late Pliocene in both South American and African lineages. Phylogenetic studies indicate ancient origins (60 Ma+) for clades characteristic of flammable sclerophyll vegetation from Australia and the Cape region of South Africa. However, as for savannas, there was a surge of speciation from the Late Miocene associated with the retreat of closed fire-intolerant forests. The wide geographic spread of increased fire activity in the last few million years suggests a global cause. However, none of the potential global factors (oxygen, rainfall seasonality, CO2, novel flammable growth forms) provides an adequate explanation as yet. The global patterns and processes of fire and flammable vegetation in the Cenozoic, especially since the Late Miocene, deserve much more attention to better understand fire in the earth system.

This study presents the first subfossil beetle (Coleoptera) records from Lithuania, from Late Glacial organic deposits. Bulk sediment samples were collected from the Pamerkiai and Zervynos Outcrops in SE Lithuania, and from the Ventė Outcrop at the eastern coast of the Curonian Lagoon, W Lithuania. Radiocarbon dating determined that the studied sediments accumulated between ~15,000–11,300 cal BP. The beetle assemblages (29–177 individuals per sample) consist of many cold-adapted species that are common from Late Glacial deposits in the British Isles, Southern Sweden, and continental Europe. True arctic species are absent from the assemblages, and it is likely that the Lithuanian beetle fauna was most similar to nearby southern regions (e.g., Poland) during the Late Glacial. Besides a variety of aquatic species and typical wetland species, many beetle species living in open environments and on sandy soils were identified. In almost all the samples, taxa associated with pine trees, willows, and birches were found, confirming previous reconstructions of a sparsely forested landscape during the climatic periods GI-1e–GI-1a (Bølling-Allerød). The species assemblages from the youngest samples, associated with GS-1 (Younger Dryas), indicate the disappearance of large aquatic macrophytes and decreasing temperatures in Southern Lithuania, but a persistence of trees in the region.

Post-disaster reconstruction offers an opportunity to address some of the fundamental causes of vulnerability that are an inherent part of mainstream housing processes located at the intersectionality of sectoral interdependencies. Well-designed initiatives in the aftermath of a disaster can help displaced populations enter a positive cycle of resilience-building using new approaches. This paper draws from a recent field study to examine the reasons for the poor performance of existing housing stock in the face of disasters and presents a chain of graded causal factors that contribute to their vulnerability. Specifically, in the context of rural housing, the paper looks at three case studies of innovative habitat reconstruction initiatives undertaken in the aftermath of major disasters in India and analyses them for their impact on building the resilience of displaced communities. The study highlights that in order to address the various causes of vulnerability of rural housing, it is important to leverage the existing connections between different dimensions of habitat development, including access to finance, choice of appropriate materials, skill-building, and safe construction methods. This is at the core of the ethos of “building back better.”

Inland waterways and their connections to marine transport systems constitute a substantial resource for the establishment of green infrastructures, flood prevention, and environmental conservation. However, these developments have numerous inherent environmental hazards such as water and air pollution, a loss of habitats, increased coastal erosion, the transfer of invasive species between connected watercourses and lakes, and the transport of pollutants through watercourses to coastal areas. Climate change may aggravate these environmental problems through changing temperatures, reduced precipitation, enhancing the adverse impact of excess nutrient discharge, and the entry of invasive species. In this study, we analyse the main European inland waterway corridors and their branches to assess the ecological viability of a pan-European inland waterway network. The environmental viability of such network depends on the right assessment of ecosystem services and protection of biodiversity. A model structure for landscape conservation, green infrastructure development, water replenishment, and ecosystem reconstruction is proposed, considering a sustainable combination of multimodal inland waterway and rail transport.

Leafhoppers (Hemiptera: Cicadellidae) form a large, diverse insect family that contributes significantly to trophic interactions and pathogen transmission in grasslands. We compared leafhopper communities of five remnant prairies and five prairie reconstructions in northeastern Kansas, USA. We hypothesized that leafhopper communities would be more conservative (i.e., greater dependence on prairies) and diverse on remnant as compared to reconstructed prairies, and that remnants would support more grass-feeders. We also predicted that sites surrounded by natural/semi-natural landscapes would have greater leafhopper diversity. We found grass feeders represented a higher proportion of leafhoppers on remnants. We found, however, no difference in mean conservatism between remnants and reconstructions, and overall levels of conservatism were lower than other grassland data sets. Mean conservatism was greater in the mid-summer than in early summer, where migratory pests such as Macrosteles were abundant. Although remnants and reconstructed prairies didn’t differ in leafhopper diversity, diversity increased with forb percent cover. The amount of natural and semi-natural land cover around sites had complex relationships with leafhopper diversity and composition. For future studies, we encourage coordinated studies on leafhopper communities across the tallgrass prairie region, with a particular focus on conservatism. Deciphering the effect of survey times, and grassland size and management, on prairie-dependent leafhoppers is important. We predict that large numbers of migratory species may lead to greater homogeneity in leafhopper communities across sites early in the season. Over time, site types and landscape features may progressively filter species and amplify local specialists, resulting in more late-season variation among sites.

The African continent is comprised of several different biomes, although savanna is the most prevalent. The current heterogeneous landscape was formed through long-term vegetation shifts as a result of the global cooling trend since the Oligocene epoch. The overwhelming trend was a shift from primarily forest, to primarily savanna. As such, faunal groups that emerged during the Paleogene/Neogene period and have species distributed in both forest and savanna habitat should show a genetic signature of the possible evolutionary impact of these biome developments. and (Colubridae) are excellent taxa to investigate the evolutionary impact of these biome developments on widespread African colubrid snakes, and whether timing and patterns of radiation are synchronous with biome reorganisation. A phylogenetic framework was used to investigate timing of lineage diversification. Phylogenetic analysis included both genera as well as other Colubridae to construct a temporal framework in order to estimate radiation times for and . Lineage diversification was estimated in Bayesian Evolutionary Analysis Sampling Trees (BEAST), using two mitochondrial markers (cyt- , ND4), one nuclear marker (c-mos), and incorporating one fossil and two biogeographical calibration points. Vegetation layers were used to classify and confirm species association with broad biome types ('closed' = forest, 'open' = savanna/other), and the ancestral habitat state for each genus was estimated. showed an ancestral state of closed habitat, but the ancestral habitat type for was equivocal. Both genera showed similar timing of lineage diversification diverging from their sister genera during the Oligocene/Miocene transition ( . 25 Mya), with subsequent species radiation in the Mid-Miocene. appeared to have undergone allopatric speciation during Mid-Miocene forest fragmentation. Habitat generalist and open habitat specialist species emerged as savanna became more prevalent, while at least two forest associated lineages within moved into Afromontane forest habitat secondarily and independently. With similar diversification times, but contrasting ancestral habitat reconstructions, we show that these genera have responded very differently to the same broad biome shifts. Differences in biogeographical patterns for the two African colubrid genera is likely an effect of distinct life-history traits, such as the arboreous habits of compared to the terrestrial lifestyle of .