Explore the vast range of titles available.

Agroforestry systems (AFS) represent combinations of trees, arable crops, and/or pastures. Being assemblages of diverse life-forms, they exhibit complex biophysical interactions. For instance, the multistrata canopies shade the understory crops by intercepting a significant amount of the incoming solar radiation. Optimizing understory productivity, thus, requires understanding the elements that affect the canopy transmittance of photosynthetically active radiation (PAR) and its spatiotemporal dynamics. We systematically reviewed the peer-reviewed literature involving 145 tropical and subtropical tree + crop combinations. The theoretical underpinnings of interspecific interactions in developing agroforestry stands were elucidated using a conceptual model. Additionally, the linkage between subcanopy PAR levels and yield was established for 11 arable crops. PAR reaching the understory and the subcanopy yield levels were tremendously variable across AFS. Relative yields ranged from 6 to 188% of the sole crops. Stage of stand development, canopy architecture, and management factors are cardinal determinants of canopy light extinction, understory PAR availability, and yield. The yield of shade-tolerant crops either increased (“over-yielding”) or remained the same as PAR levels decreased within certain limits, albeit with intraspecific variations. The tree-crop interaction effects on yield were positive, negative, or neutral. In total, 19 cases showed positive responses, 29 were neutral, and 113 were negative, with a few overlapping responses depending on the tree, crop, and management. This implies that the key to ecological intensification is component selection and management. Agroforestry, while containing the loss of, maintaining, or even increasing understory yields, thus maximizes overall (tree + crop) outputs and land equivalent ratio.

Questions: What are the stand development patterns and successional processes occurring in a Quercus petraea-Fagus sylvatica old-growth forest? Do these two sympatric species have different recruitment and canopy accession strategies? How do they respond to disturbance? Location: Old-growth Q. petraea-F. sylvatica forest in Runcu Grosi Natural Reserve, Romania. Methods: Increment cores were taken from 110 similarly sized pairs of standing dead and living sessile oak trees, and additionally from the 1 10 most competitive neighbouring living beech trees, growing in the best-preserved part of the reserve. Release events were detected using the boundary line method. Recruitment and canopy accession strategies were defined using tree juvenile growth rate, overall growth patterns and the presence or absence of a major crown release. Results: Sessile oak establishment occurred between 1800 and 1859, while beech have regenerated continuously, with greater in-growth after 1860. Tree recruitment followed species-specific requirements, with most sessile oak having a gap origin (82%) and many beech regenerating in the understorey (41 %). Release events occurred continuously during the period analysed (1820-2003), but the percentage of trees per decade that showed releases was < 15%. A larger number of releases were recorded in beech compared to sessile oak. Living and dead sessile oak originated under similar conditions, but after the 1900s they experienced different canopy disturbances regimes. Conclusions: The investigated stand appears to originate after a large-scale disturbance that promoted sessile oak recruitment. Thereafter, regular small-scale disturbances favoured beech, which became a competitor to sessile oak and achieved faster growth despite a less favourable initial canopy position. Mortality started to occur in oak. Stand maturation and increasing competition could favour the occurrence of large-scale windthrow, which would again promote sessile oak.

This study aimed to compare the effects of whole-tree and stem-only harvesting in hemiboreal forests in Latvia. Chemistry of soil solution, precipitation, litter and needles, as well as tree parameters in regenerated stands were measured from 2012 to 2021 in oligotrophic and mesotrophic Scots pine sites with mineral soils and a eutrophic Norway spruce site with drained peat soil. Compared to the undisturbed control, the soil solution pH was significantly lower and N-NO3− and K+ concentrations were significantly higher after stem-only harvesting at the oligotrophic site, stem-only and whole-tree harvesting at the mesotrophic site and whole-tree harvesting at the eutrophic site. The height growth of the regenerated stands in all sites was similar for both harvesting methods six years after planting. More slender trees were observed after whole-tree harvesting than after stem-only harvesting at the eutrophic site. Whole-tree harvesting produced significant negative short-term effects on height growth in the oligotrophic site four to six years after harvest. The nutrient levels of needles differed with harvest intensity but did not indicate insufficient nutrient availability with any type of harvesting.

• Global warming is expected to exacerbate the duration and intensity of droughts in the western United States, which may lead to increased tree mortality. A prevailing proximal mechanism of drought-induced tree mortality is hydraulic damage, but predicting tree mortality from hydraulic theory and climate data still remains a major scientific challenge. • We used forest inventory data and a plant hydraulic model (HM) to address three questions: can we capture regional patterns of drought-induced tree mortality with HM-predicted damage thresholds; do HM metrics improve predictions of mortality across broad spatial areas; and what are the dominant controls of forest mortality when considering stand characteristics, climate metrics, and simulated hydraulic stress? • We found that the amount of variance explained by models predicting mortality was limited (R² median = 0.10, R² range: 0.00–0.52). HM outputs, including hydraulic damage and carbon assimilation diagnostics, moderately improve mortality prediction across the western US compared with models using stand and climate predictors alone. • Among factors considered, metrics of stand density and tree size tended to be some of the most critical factors explaining mortality, probably highlighting the important roles of structural overshoot, stand development, and biotic agent host selection and outbreaks in mortality patterns.

1. Mixed species forests can often be more productive and deliver higher levels of ecosystem services and functions than monocultures. However, complementarity effects for any given tree species are difficult to generalize because they can vary greatly along gradients of climatic conditions and resource availability. Identifying the conditions where species diversity can positively influence productivity is crucial. To date, few studies have examined how growth complementarity across species and mixture types is modulated by stand and environmental factors, and fewer have considered more than one or two factors. 2. We investigated how complementarity effects for several major Central European tree species change with climatic and edaphic conditions, and with stand structural characteristics, including species composition. We used data from the Swiss National Forest Inventory, which is based on 3,231 plots of pure and mixed stands (19 mixture types) across a broad environmental gradient, to test (i) how mixing effects change depending on the identity of the admixed species and (ii) if complementarity consistently increases when environmental conditions become harsher. 3. The magnitude, whether positive or negative, of complementarity increased with increasing stand density and stand developmental stage, but no general pattern could be identified across mixture types. Complementarity for many species increased as drought intensity and temperature increased, but not for all species and mixture types. While soil conditions, nitrogen and site topography influenced complementarity for many species, there was no general pattern (increases and decreases were observed). 4. Synthesis. Our study indicates that complementarity varies strongly with stand density and stand development as well as with topographic, climatic and soil conditions. This emphasizes the need to account for site-dependent conditions when exploring mixture effects in relation to forest productivity. We found that under certain conditions (i.e. increasing drought, higher temperature), mixed forests can promote individual tree growth in Central European temperate forests. However, careful assessments depending on the species composing the stands are required under changing resource availability as well as under different levels of stand density and development.

Key messageThe proposed height–diameter model applicable to many tree species in the multi-layered and mixed stands across Czech Republic shows a high accuracy in the height prediction. This model can be useful for estimating forest yield and biomass, and simulation of the vertical stand structures.We developed a generalized nonlinear mixed-effects height–diameter (H–D) model applicable to Norway spruce (Picea abies (L.) Karst.), European beech (Fagus sylvatica L.) and other conifer and broadleaved tree species using the modelling method that includes dummy variables accounting for species-specific height differences and random component accounting for within- and between-sample plot height differences and randomness in the data. We used two large datasets: the first set (model fitting dataset) originated from permanent research sample plots and second set (model-testing dataset) originated from the Czech national forest inventory (NFI) sample plots. The former dataset comprises 224 sample plots with 29,390 trees and the latter dataset comprises 14,903 sample plots with 382,540 trees, each representing wide variabilities of tree size, ecological zone, growth condition, stand structure and development stage, and management regime across the country. Among the four versatile growth functions evaluated as base functions with diameter at breast height (DBH) included as a single predictor, the Chapman-Richards function showed the most attractive fit statistics. This function was then extended through the integration of other predictor variables, which better describe the stand density (stand basal area), stand development and site quality (dominant height), competition (ratio of DBH to quadratic mean DBH), that would act as modifiers of the original parameters of the function. The mixed-effects H–D model described a large part of the variations in the H–D relationships (\\[R_adj^2\\] = 0.9182; RMSE = 2.7786) without substantial trends in the residuals. Testing this model against model-testing dataset confirmed the model’s high accuracy. The model may be used for estimating forest yield and biomass, and therefore will serve as an important tool for decision making in forestry.

Reducing tree densities through silvicultural thinning has been widely advocated as a strategy for enhancing resistance and resilience to drought, yet few empirical evaluations of this approach exist. We examined detailed dendrochronological data from a long-term (>50 years) replicated thinning experiment to determine if density reductions conferred greater resistance and/or resilience to droughts, assessed by the magnitude of stand-level growth reductions. Our results suggest that thinning generally enhanced drought resistance and resilience; however, this relationship showed a pronounced reversal over time in stands maintained at lower tree densities. Specifically, lower-density stands exhibited greater resistance and resilience at younger ages (49 years), yet exhibited lower resistance and resilience at older ages (76 years), relative to higher-density stands. We attribute this reversal to significantly greater tree sizes attained within the lower-density stands through stand development, which in turn increased tree-level water demand during the later droughts. Results from response-function analyses indicate that thinning altered growth-climate relationships, such that higher-density stands were more sensitive to growing-season precipitation relative to lower-density stands. These results confirm the potential of density management to moderate drought impacts on growth, and they highlight the importance of accounting for stand structure when predicting climate-change impacts to forests.

1. Understorey vegetation comprises the majority of species diversity and contributes greatly to ecosystem functioning in boreal forests. Although patterns of understorey abundance, species diversity and composition associated with forest stand development are well researched, mechanisms driving these patterns remain largely speculative. 2. We sampled fire-origin stands of varying stand ages and overstorey compositions on mesic sites of the boreal forest of Canada and used structural equation modelling (SEM) to link time since fire (stand age), light availability and heterogeneity, substrate heterogeneity and soil nitrogen to understorey vegetation cover and species diversity. 3. The most parsimonious model for total understorey cover showed a positive direct effect of stand age (r = .43) and an indirect effect via mean light level (0.18) and shrub cover (-0.11), with a positive total effect (0.50); the per cent broadleaf canopy had a direct negative effect (-0.22) and an indirect effect via shrub cover (-0.11). The model for total understorey species richness showed an indirect effect of stand age via mean light (0.24), light heterogeneity (0.10) and substrate heterogeneity (0.07), with a positive total effect (0.52); per cent broadleaf canopy had an indirect effect via light heterogeneity (0.09), and substrate heterogeneity (-0.10). Soil nitrogen did not significantly influence either understorey cover or species richness. The models for vascular plants followed similar trends to those for total understorey cover and species richness; however, there was an opposite indirect effect of light heterogeneity for both cover and species richness of non-vascular plants. Shrub cover had positive direct and negative direct and indirect effects on both vascular and non-vascular cover and species richness. 4. Synthesis. Our findings indicate that understorey cover and species diversity are driven by time since disturbance, light availability as influenced by overstorey and shrub layers, but with important additional effects mediated by light and substrate heterogeneity. Non-vascular understorey vegetation is more strongly determined by time since disturbance than vascular vegetation, and negatively affected by broadleaf tree abundance. The overall results highlight the importance of colonization, light availability and heterogeneity, substrate specialization and growth dynamics in determining successional patterns of boreal forest understorey vegetation.

Background and aims Soil phosphorus (P) can often regulate plant productivity. However, the medium- to long-term effects of reforestation on P cycle within the continuum of plant, litter, and soil (including deep soil layers), and the subsequent regulation of soil P storage and its fractions, remain unclear. Methods We determined soil (0–100 cm) P fractions, acid phosphatase, stoichiometric ratios, root and leaf N/P ratios, leaf P resorption efficiency (PRE), and leaf-litter P concentration of 32-, 45-, and 60-year-old Pinus massoniana reforestation in southwest China. Results The storage of soil labile, moderately labile, and occluded P decreased with the increase of stand age. The concentration of NaHCO 3 -P i , NaHCO 3 -P o , and total labile P in top-soil layers decreased with the increase of stand age. The concentration of NaOH-P o and total moderately labile P in the entire soil profile declined over time. The concentration of C.HCl-P i and total occluded P in the 0–100 cm soil layer were lower in the 60-year-old stand than in the 32-year-old stand. The PRE, leaf N/P ratio, top-soil C/P ratio and acid phosphatase activity increased, and the leaf-litter P concentration decreased with the increase of stand age. Conclusions P. massoniana trees secreted more acid phosphatase and increased PRE to compensate for the decrease in soil P availability with stand development, which in turn decreased leaf-litter P concentration and thus resulted in a depletion of soil P. Overall, our results highlighted that P limitation of P. massoniana reforestation increased with stand development.

Climate change alters forest development pathways, with consequences for ecosystem services and biodiversity. As the rate of warming increases, ecosystem change is expected to accelerate. However, ecosystem dynamics can have many causes unrelated to climate (for example, disturbance and stand development legacies). The compound effects of multiple drivers remain largely unclear. Here, we assessed forest dynamics over 28 years at Berchtesgaden National Park (BGNP), Germany, quantifying the spatiotemporal patterns and unraveling the drivers of forest change. We analyzed high-density forest inventory data, consisting of three consecutive censuses of 3759 permanent sample plots (132,866 tree records in total). We used semi-variograms to analyze spatial patterns of change, and boosted regression trees to quantify the effect of 30 covariates on changes in nine indicators of forest structure and composition. Over the 28 years investigated, the forests of BGNP were becoming denser, structurally more complex, and more species rich. Changes in forest structure were more pronounced and spatially correlated on the landscape than changes in tree species composition. Change rates of all indicators increased over time, signifying an acceleration of forest dynamics since the 1980s. Legacies and climate were the most important drivers of change, but had diverging impacts. Although forest change accelerated with increasing temperature, high legacy levels typical for late development stages dampened it. We here provide evidence for accelerating forest dynamics in mountain forests of the Alps, with potentially far-reaching consequences for biodiversity and ecosystem processes. We highlight that unmanaged forest development toward old-growth conditions could counteract climatemediated acceleration of forest change.