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In antiquity Archilochus of Paros was considered a poet rivalled only by Homer and Hesiod, yet he has been relatively neglected by modern scholarship. This is largely due to the fragmentary state of his surviving poetry, though our knowledge has expanded significantly since the middle of the twentieth century as new papyrological finds continue to augment the corpus and our understanding of the poet and his work evolves. This volume is the first ever complete commentary on Archilochus, filling a substantial gap in scholarship on archaic Greek poetry and playing an important and timely role in re-establishing him as a major author and in locating the recent discoveries in the broader context of his oeuvre. Presenting the fragmentary texts alongside brand new translations, the volume also contains a comprehensive introduction offering an accessible guide to Archilochus' work and context, and a detailed commentary providing textual, literary, and historical analysis of all of his surviving poetry and discussing broader questions of performance and genre in early Greek poetic culture. The scope and depth of the analysis not only highlights the diversity and sophistication of Archilochus' work, but also sheds new light on our understanding of Greek iambus and elegy, while his influence on later writers means that the commentary will be of significance to scholars and students of Hellenistic and Roman literature, and the later lyric tradition, as well as archaic and classical Greek literature.

Background The study of speciation has expanded with the increasing availability and affordability of high-resolution genomic data. How the genome evolves throughout the process of divergence and which regions of the genome are responsible for causing and maintaining that divergence have been central questions in recent work. Here, we use three pairs of species from the recently diverged bee hummingbird clade to investigate differences in the genome at different stages of speciation, using divergence times as a proxy for the speciation continuum. Results Population measures of relative differentiation between hybridizing species reveal that different chromosome types diverge at different stages of speciation. Using FST as our relative measure of differentiation we found that the sex chromosome shows signs of divergence early in speciation. Next, small autosomes (microchromosomes) accumulate highly diverged genomic regions, while the large autosomes (macrochromosomes) accumulate genomic regions of divergence at a later stage of speciation. Conclusions Our finding that genomic windows of elevated FST accumulate on small autosomes earlier in speciation than on larger autosomes is counter to the prediction that FST increases with size of chromosome (i.e. with decreased recombination rate), and is not represented when weighted average FST per chromosome is compared with chromosome size. The results of this study suggest that multiple chromosome characteristics such as recombination rate and gene density combine to influence the genomic locations of signatures of divergence.

\"This is the first book to study the impact of invective poetics associated with early Greek iambic poetry on Roman imperial authors and audiences. It demonstrates how authors as varied as Ovid and Gregory Nazianzen wove recognizable elements of the iambic tradition (e.g. meter, motifs, or poetic biographies) into other literary forms (e.g. elegy, oratorical prose, anthologies of fables), and it shows that the humorous, scurrilous, efficacious aggression of Archilochus continued to facilitate negotiations of power and social relations long after Horace's Epodes. The eclectic approach encompasses Greek and Latin, prose and poetry, and exploratory interludes appended to each chapter help to open four centuries of later classical literature to wider debates about the function, propriety and value of the lowest and most debated poetic form from archaic Greece. Each chapter presents a unique variation on how these imperial authors became Archilochus - however briefly and to whatever end\"-- Provided by publisher.

For a second decade, from November 2008 to March 2018, we examined the species diversity, return rate, longevity, and migration routes of hummingbirds overwintering in southern Alabama and the panhandle, northern peninsula, and central peninsula of Florida. We captured and banded 11 species of hummingbirds (n= 1,870 individuals), including 931 Rufous Hummingbirds (Selasphorus rufus; 49%), 583 Ruby-throated Hummingbirds (Archilochus colubris; 31%), 219 Black-chinned Hummingbirds (A. alexandri; 12%), 49 Buff-bellied Hummingbirds (Amazilia yucatanensis), 49 Calliope Hummingbirds (S. calliope), 19 Allen's Hummingbirds (S. sasin), 11 Broad-tailed Hummingbirds (S. platycercus), 6 Broad-billed Hummingbirds (Cynanthus latriostris), 1 Anna's Hummingbird (Calypte anna), 1 Costa's Hummingbird (Calypte costae), and 1 White-eared Hummingbird (Hylocharis leucotis). Site fidelity was high, with 299 hummingbirds of 5 species returning at least once. Two Rufous Hummingbirds migrated to our study area in the autumn from the West (Oklahoma and Texas). One Ruby-throated Hummingbird banded in Lakeland, Florida, was recovered in Utopia, New Brunswick, Canada, suggesting the overwintering Ruby-throated Hummingbirds in our study area are migratory and not sedentary.

Neonicotinoids are neurotoxic systemic insecticides applied extensively worldwide. The impacts of common neonicotinoids like imidacloprid on non-target invertebrate pollinators have been widely studied, however effects on vertebrate pollinators have received little attention. Here, we describe the first study evaluating the effects of short-term (3 d) exposure to a range of environmentally relevant concentrations ( 0.2 μ g g - 1 to 2.5 μ g g - 1 · Body Weight) of imidacloprid on wild-caught ruby-throated hummingbirds. Within 2 h of exposure, hummingbirds exhibited a significant depression in energy expenditure (up to 25 % ± 11 % ). We did not observe significant effects on foraging behaviour measured in the subsequent 2 h to 4 h, although the effect size estimate was large (0.29). We also analyzed tissues collected 24 h after the final dose and did not observe significant effects on immune response or cholinesterase activity, although this may be related to our small sample size. We determined that hummingbirds excrete imidacloprid quickly (elimination half-life of 2.1 h ± 0.1 h ) relative to other bird species. Hummingbirds have high energetic demands and store relatively little energy, especially during migration and breeding seasons. Therefore, changes in their metabolism following exposures to imidacloprid observed herein could bear important survivorship consequences for hummingbirds.

Although the breeding ranges of Archilochus alexandri (Black-chinned Hummingbird) and Archilochus colubris (Ruby-throated Hummingbird) are narrowly parapatric in central Texas and central and southern Oklahoma, there have been few reports of hybridization in the literature and no well-documented hybrid specimens. Here we provide a comprehensive assessment of two male hybrids collected, respectively, in Grayson County, Texas, and East Baton Rouge Parish, Louisiana. As has been the pattern in other hummingbird hybrids, both specimens exhibit a blended mosaic of plumage characters of the parental species. Sequence for mitochondrial cytochrome c oxidase subunit 1 (CO1) for the hybrid specimen from East Baton Rouge Parish (LSU 182,831) confirms A. colubris as the female parent.

Many plants attract and reward pollinators with floral scents and nectar, respectively, but these traits can also incur fitness costs as they also attract herbivores. This dilemma, common to most flowering plants, could be solved by not producing nectar and/or scent, thereby cheating pollinators. Both nectar and scent are highly variable in native populations of coyote tobacco, Nicotiana attenuata, with some producing no nectar at all, uncorrelated with the tobacco's main floral attractant, benzylacetone. By silencing benzylacetone biosynthesis and nectar production in all combinations by RNAi, we experimentally uncouple these floral rewards/attractrants and measure their costs/benefits in the plant's native habitat and experimental tents. Both scent and nectar increase outcrossing rates for three, separately tested, pollinators and both traits increase oviposition by a hawkmoth herbivore, with nectar being more influential than scent. These results underscore that it makes little sense to study floral traits as if they only mediated pollination services. Flowering plants have evolved a number of different approaches to reproduction. Some use their own pollen and self-fertilize, while others use pollen from other nearby plants. This fertilization by other plants is called ‘outcrossing’ and introduces new genetic variation into each generation, which is extremely important for the evolutionary process. Some flowering plants rely on animals to help with pollination, attracting visitors with floral scents and rewarding the visitors with sugar-rich nectar. But scent and nectar also attract herbivores that damage the plants. This causes a dilemma for flowering plants, which has led some to evolve to not produce scent or to offer no nectar while masquerading as a plant that does. Previous studies into the costs and benefits of such strategies have looked at the effects of either floral scent or nectar, but no-one has uncoupled the effects of these two traits on both pollination and herbivore attack. Kessler et al. have addressed this issue in wild tobacco plants, which can both self-fertilize and outcross, and which produce varying amounts of scent and nectar. The experiments were conducted under mesh tents and in field trials in the plant's natural habitat: the Great Basin Desert in Utah. Kessler et al. used a gene-silencing technique called ‘RNA interference’ to inhibit the production of scent or nectar, either separately or together. When grown in field trials, under conditions that prevent self-fertilization, these tobacco plants are frequently visited by a hummingbird and three species of hawkmoth. All four of these animals pollinate the tobacco plants, but one of the moths also lays eggs that hatch into caterpillars, which damage the plant. Kessler et al. monitored the effects that the loss of scent, nectar or both had on visits by each pollinator and on outcrossing. These experiments revealed that scent is essential to attract one hawkmoth species but not for another (called Hyles lineata). Furthermore, while, the hummingbird needs nectar, the H. lineata moth does not; but this moth won't visit flowers that lack both scent and nectar. The experiments also show that, for the moth that lays its eggs on the tobacco plants, both scent and nectar increase pollination and egg laying, but nectar has a stronger effect. Thus reducing nectar, as this tobacco plant does in the wild, is one strategy that can be used to reduce herbivore attack by caterpillars. Together, these findings highlight that it is important to study both herbivores and pollinators when attempting to understand the evolution of floral traits.

Hummingbirds are vital members of terrestrial ecosystems, and because of their high metabolic requirements, they serve as indicators of ecosystem health. Monitoring the parasitic infections of hummingbirds is thus especially important. Haemosporidians, a widespread group of avian blood parasites, are known to infect hummingbirds, but little is known about the prevalence and diversity of these parasites in hummingbirds. The prevalence of haemosporidians in several hummingbird species was examined and we compared 4 different tissue types in detecting parasites by polymerase chain reaction (PCR). Blood samples from 339 individuals of 3 different hummingbird species were tested, and 4 individuals were found positive for haemosporidian infection, a prevalence of 1.2%. Hummingbird carcasses (n = 70) from 5 different hummingbird species were also sampled to assess differences in detection success of haemosporidians in heart, kidney, liver, and pectoral muscle tissue samples. Detection success was similar among tissue types, with haemosporidian prevalence of 9.96% in heart tissue, 9.52% in kidney tissue, 10.76% in liver tissue, and 11.76% in pectoral muscle tissue. All tissue samples positive for haemosporidian infection were from the Black-chinned Hummingbird (Archilochus alexandri). Possible reasons for low prevalence of these blood parasites could include low susceptibility to insect vectors or parasite incompatibility in these hummingbirds.

External feather rinses and homogenized whole-carcass tissue matrix from two hummingbird species found in California ( Calypte anna and Archilochus alexandri ) were analyzed for the presence of nine insecticides commonly used in urban settings. Using a liquid chromatography-high-resolution mass spectrometry (LC-HRMS) analytical method, samples were quantitatively tested for the following neonicotinoids: dinotefuran, nitenpyram, thiamethoxam, acetamiprid, thiacloprid, clothianidin, imidacloprid, and sulfoxaflor. This analytical method was also used to qualitatively screen for the presence of approximately 150 other pesticides, drugs, and natural products. Feather rinsates from both hummingbird species had detectable concentrations of carbamate and neonicotinoid classes of insecticides. Combined results of the rinsate and homogenized samples ( n = 64 individual hummingbirds) showed that 44 individuals (68.75%) were positive for one to four target compounds. This study documented that hummingbirds found in California are exposed to insecticides. Furthermore, feather rinsates and carcass homogenates are matrices that can be used for assessing pesticide exposure in small bird species. The small body size of hummingbirds limits traditional sampling methods for tissues and whole blood to evaluate for pesticide exposure. Thus, utilization of this analytical method may facilitate future research on small-sized avian species, provide insight into pesticide exposure, and ultimately lead to improved conservation of hummingbirds.

Haemosporidian blood parasites are transmitted to a wide range of avian hosts via blood-sucking dipteran vectors. Microscopy has revealed an impressive diversity of avian haemosporidia with more than 250 species described. Moreover, PCR and subsequent sequence analyses have suggested a much greater diversity of haemosporidia than morphological analyses alone. Given the importance of these parasites, very few studies have focused on the charismatic hummingbirds. To date, three Haemoproteus species ( Haemoproteus archilochus , Haemoproteus trochili , and Haemoproteus witti ) and one Leucocytozoon species ( Leucocytozoon quynzae ) have been described in blood samples taken from hummingbirds (Trochilidae). Unconfirmed Plasmodium lineages have also been detected in hummingbirds. Here, we report the detection of H. archilochus in two hummingbird species ( Calypte anna and Archilochus alexandri ) sampled in Northern California and perform a phylogenetic analysis of mitochondrial cytochrome b (cyt b ) gene lineages. A total of 261 hummingbirds (157 C. anna , 104 A. alexandri ) were sampled and screened for blood parasites using PCR and microscopy techniques. Combining both methods, 4 (2.55%) haemosporidian infections were detected in C. anna and 18 (17.31%) haemosporidian infections were detected in A. alexandri . Molecular analyses revealed four distinct H. archilocus cyt b lineages, which clustered as a monophyletic clade. No species of Plasmodium or Leucocytozoon were detected in this study, raising the possibility of specific vector associations with hummingbirds . These results provide resources for future studies of haemosporidian prevalence, diversity, and pathogenicity in California hummingbird populations.