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Interactions between flowering time genes were examined in a doubled haploid barley (Hordeum vulgare) population segregating for H. vulgare VERNALIZATION1 (HvVRN1), HvVRN2, and PHOTOPERIOD1 (PPD-H1). A deletion allele of HvVRN2 was associated with rapid inflorescence initiation and early flowering, but only in lines with an active allele of PPD-H1. In these lines, the floral promoter FLOWERING LOCUS T (HvFT1) was expressed at high levels without vernalization, and this preceded induction of HvVRN1. Lines with the deletion allele of HvVRN2 and the inactive ppd-H1 allele did not undergo rapid inflorescence initiation and were late flowering. These data suggest that HvVRN2 counteracts PPD-H1 to prevent flowering prior to vernalization. An allele of HvVRN1 that is expressed at high basal levels (HvVRN1-1) was associated with rapid inflorescence initiation regardless of HvVRN2 or PPD-H1 genotype. HvFT1 was expressed without vernalization in lines with the HvVRN1-1 allele and HvFT1 transcript levels were highest in lines with the active PPD-H1 allele; this correlated with rapid apex development postinflorescence initiation. Thus, expression of HvVRN1 promotes inflorescence initiation and up-regulates HvFT1. Analysis of HvVRN1 expression in different genetic backgrounds postvernalization showed that HvVRN2, HvFT1, and PPD-H1 are unlikely to play a role in low-temperature induction of HvVRN1. In a vernalization responsive barley, HvFT1 is not induced by low temperatures alone, but can be induced by long days following prolonged low-temperature treatment. We conclude that low-temperature and daylength flowering-response pathways are integrated to control expression of HvFT1 in barley, and that this might occur through regulation of HvVRN2 activity.

People's understandings of natives and newcomers in early America usually proceed from two problematic assumptions: landlocked Indians and European seas. Here, Bahar invites people to reconsider these constructions with his exploration of an Atlantic-oriented Indian people who systematically plundered the waves of European ships, sailors, and cargo. For two centuries Indian marine-warriors managed and manipulated colonialism by forging an extractive subsidiary economy from a European Atlantic commercial nexus. This story remains forgotten, he argues, because Atlantic world history and American Indian history have been two ships passing in the night, fostering a historiographical dissonance undergirded by popular stereotypes about Indians.

We argue in favor of merely disjunctive effects, namely cases in which an event or fact, C, is not a cause of an effect, E₁, and is also not a cause of a distinct effect, E₂, and yet C is a cause of the disjunctive effect (E₁ or E₂). Disjunctive effects let us retain the additivity and the distributivity of causation. According to additivity, if C is a cause of E₁ and C is a cause of E₂, then C is a cause of E₁ and E₂. According to distributivity, if C is a cause of E₁ and E₂, then C is a cause of E₁ and C is a cause of E₂. We draw an analogy between causation and intensional notions like believing, wanting, and owing, which also admit of merely disjunctive cases. We argue that both the Lewisian counterfactual account of causation (including its recent emendation by Sartorio) and the contrastive account of causation fail to properly account for this phenomenon.

Improved boron (B) tolerance has been an objective of barley breeding programs in regions where B toxicity occurs. Traits associated with B tolerance have been mapped on chromosomes 2H and 4H and it has been proposed that these be used for marker assisted selection for B tolerance. However, there has been little or no improvement in yield using this strategy. This study examined the reasons for the small yield differences among different lines of barley that differ in B tolerance. Experiments used backcross lines derived from crosses between the B-tolerant landrace Sahara 3771 and two adapted recurrent parents, Sloop and VB9104. Lines with different combinations of the Sahara 3771 alleles on chromosomes 2H and 4H were grown over three growing seasons at sites where barley is prone to B toxicity. Grain yields of the backcross lines were similar to or lower than those of the recurrent parents despite showing differences in the expression of B toxicity symptoms and in B concentration in vegetative tissue. There were few significant differences in grain yield among the backcross lines. Variation in dry matter production among the backcross lines in each of the three growing seasons was unrelated to shoot B concentrations while grain yield was correlated with shoot B concentration only among the backcross lines of VB9104 in one season. In this case the yield loss was 4% per 10 mg kg⁻¹ increase in shoot B concentration. Variation in shoot B concentration and yield across seasons was much greater than that observed among the different barley lines. Reduced B accumulation was associated with higher shoot sodium concentration among the Sloop backcross lines. The results suggest that yield gains from selection based largely on B exclusion and symptoms expression may be small and strongly affected by site and seasonal effects. In the regions where other soil constraints, such as soil salinity and micronutrient deficiencies are also important, reducing B uptake alone may have little effect on yield if these other soil properties are also limiting yields.

A short story is presented.

In the 18th century, Bermuda depended on the sea for its economic survival. Its merchant fleet helped to carry the trade of Britain's expanding empire and was rewarded with the food, timber and cash needed to sustain the island's inhabitants, shipbuilding industry and economy. The success of the carrying trade rested firmly on the shoulders of Bermuda's white and black mariners, who shared the labor--but unequally reaped the benefits--of their work.